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British Museum (Natural History) 

Ruwenzori Expedition 


Volume II, Number 12 

If; ' i> y. ^i§I§ll 



Trustees of the British Museum (Natural History) 
Issued October, 1965 Price £1 12*. od. 

© Trustees of the British Museum (Natural History) 1965 


Micr olepidop tera 

By J. D. Bradley 
Commonwealth Institute of Entomology 

The collection of Microlepidoptera formed by Mr. D. S. Fletcher during the Ruwenzori 
Expedition 1952-3 is among the most important of this group known from the highlands of 
East Africa. The special attention paid by the Expedition to the upper limits of the montane 
rain forest zone and the ericaceous and alpine belts lying between 8,500 ft. and 14,000 ft., where 
hitherto little collecting of Microlepidoptera appears to have been done, has yielded much new 
information concerning the composition and affinities of the microlepidopterous fauna of the 
Ruwenzori Range. Many of the species taken at these higher altitudes have proved to be either 
new to science or species previously known oidy from type material. The fauna of the regions 
below the level of the montane forest zone, which begins at about 5,000 ft., is more typically 
African in character and includes species with a wider distribution. 

The total number of specimens collected by Mr. Fletcher belonging to the families generally 
classed as Microlepidoptera, excluding Pyraloidea, exceeds 1,200. A little over half of these, 
mainly specimens from the higher altitudes, have now been studied and the results are presented 
in this paper. Such material from the lower levels as has so far been studied is included: the 
remainder still awaits study. 

The material examined has also included some from other sources, notably from the collections 
made by F. W. Edwards and D. R. Buxton during the British Museum Expedition to East 
Africa in 1934-5, an d by the G. F. de Witte Expedition to the Congo in 1947. For the privilege 
of examining the Microlepidoptera collected by the de Witte Expedition I am indebted to Dr. 
A. Houben, Director of the Institut des Pares Nationaux, Brussels, and his predecessor the late 
Dr. V. van Straelen. Only the very small part of this extensive collection from the Congo which 
supplements the Ruwenzori collection is studied in this paper. 

The material discussed in the present report includes ninety species, forty-eight of them new 
to science, and a new subspecies. No new genera are erected, species being placed in existing 
genera, though in a number of cases, it is evident that the assignment is taxonomically not 
strictly tenable. Where such provisional generic combinations occur I have commented on them 
in the text. This course has been adopted because extensive generic revision is required in prac- 
tically every family mentioned herein, and it is considered inappropriate to attempt such revision 
in the present study. 

The types of new species described are in the British Museum (Natural History) unless indicated 
otherwise. Colour terms used in the descriptions are based on Ridgway (1912) and KLornerup 
& Wanscher (1963). 



Ruwenzori Expedition, 1952. Volume II, Number 12 

A detailed list of collecting localities and the itinerary of the Expedition appears in an earlier 
part under the series title (Evans & Fletcher, 1958). The geographical distribution of previously 
described species recorded below is given so far as known. 

The illustrations of the wings (Figs. 1-62) are reproduced from photographs taken by Mr. P. 
Green, a staff photographer of the British Museum (Natural History). As the individual illustra- 
tions are considerably enlarged and of varying magnifications, the wing expanse (from wing tip 
to wing tip) of each species has been given, in parentheses, after the caption. The photomicro- 
graphs of the genitalia (Figs. 63-215) were taken by Mr. J. V. Brown, also of the museum 
photographic section, and I am indebted both to him and his colleague for these valuable visual 
aids to identification. 

The study of Mr. Fletcher's Ruwenzori material began some years ago, while I was on the 
staff of the British Museum (Natural History), and has continued intermittently. It gives me 
pleasure to acknowledge the encouragement that I received initially from Mr. N. D. Riley, c.b.e., 
then Keeper of Entomology, and subsequently from his successors Dr. W. E. China, c.b.e. and 
Mr. J. P. Doncaster ; and also from Mr. W. H. T. Tarns, to whom I am additionally indebted 
for reading and criticising the original manuscript. I am also greatly obliged to Dr. A. Diakonoff, 
of the Rijksmuseum van Natuurlijke Historie, Leiden, for his opinion regarding the generic 
affinities of certain of the Tortricidae. 


Evans, G. O. & Fletcher, D. S., 1958. Introduction and List of Localities. Ruwenzori Expedition 1952, 
1 : iii-vii, 1 pi., 1 map. 

Kornerup, A. & Wanscher, J. H., 1963. Handbook of Colour, pp. 1-224, incl. 30 col. pis. Methuen & 
Co. Ltd., London. English translation. 

Ridgway, R., 1912. Color Standards and Color Nomenclature, pp. 1-44, 53 col. pis. Washington, D.C. 


Euxanthis limenias Mcyrick 
Euxanthis limenias Meyrick, 1928, Exot. Microlepidopt., 3 : 437. 
ruwenzori: Mahoma River, 6,700 ft. (Fletcher), 2 6*. 

The two Ruwenzori specimens show evidence of variation within this species, previously 
known only from the unique type male from the Congo. Unlike the type they exhibit little trace 
of reddish brown suffusion in the coloration of the fore wing, and the wing markings are a 
comparatively blackish brown; in addition the semioval whitish patch on the costa is suffused 
with greyish and streaked with fine blackish lines. A prominent feature is the broad sinuous 
whitish band, suffused with greyish, extending from near the base of the fore wing to the middle 
of the inner margin (dorsum), curving apicad and finally merging with the distal margin of the 
semioval costal patch; in the type this band is almost completely obliterated by a brownish 
suffusion except for a short whitish furcation emitted from the costal patch. 

The genitalia of the Ruwenzori specimens have been compared with those of the type and 
found to be identical except for slight differences in the cornuti of the aedeagus. In the type all 

Microlepidoptera 83 

three cornuti - one very small thorn-like cornutus and two long slender cornuti - are straight, 
while in the Ruwenzori specimens the two larger cornuti are strongly curved. This difference at 
the most probably indicates no more than subspeciation. 
Distribution: Congo. 

Trachybyrsis euglypta Meyrick 

Trachybyrsis euglypta Meyrick, 1927, Exot. Microlepidopt.. 3 : 368. 

ruwenzori: Mahoma River, 6,700 ft. (Fletcher), 1 Nyinabitaba, 8,650 ft. (Fletcher), 1 
Nyamaleju, 10,530 ft. (Fletcher), 1 Bigo, 11,400 ft. (Fletcher), 1 q\ 

This species has hitherto been known only from the type, a female erroneously described by 
Meyrick as a male. The four Ruwenzori specimens, collected at various altitudes between 6,000 
ft. and 12,000 ft., show that the species has a vertical distribution which includes rain forest and 
the ericaceous and alpine belts. Of interest is that the wing expanse of the moths increases with the 
altitude of the localities. The Mahoma River specimen, taken at 6,700 ft., is the smallest, with a 
wing span of 24 mm.; the Nyinabitaba specimen, 8,650 ft., has a wing span of 30 mm., which 
is similar to that of the type from Niansa, 5-6,500 ft.; the Nyamaleju specimen, 10,530 ft., 
measures 32 mm.; and the Bigo specimen, 11,400 ft., is the largest and measures 35 mm. 

The Ruwenzori specimens differ superficially from the type in having the fore wing distinctly 
more greenish in colour. 

Distribution: Rwanda. 

Trachybyrsis hypsitropha sp.n. (Figures 1, 63) 

o* 37 mm. Labial palpus warm buff, darkened with fuscous irroration anteriorly. Head warm 
buff mixed with dark ochraceous-buff especially on crown. Thorax and tegula ochraccous-buff 
mixed with ochraceous-tawny, sometimes suffused greyish. Antenna with scape dark ochraceous- 
buff above, whitish below; flagellum warm buff, underside suffused with greyish and with basal 
segments marked with fuscous. Fore wing ground coloration light buff or light ochraceous-buff, 
almost entirely overlaid with fuscous suffusion liberally sprinkled with blackish, the whole 
sparsely irrorate with isabclla color, giving a partially greenish tinge to the wing; markings 
diffuse and ill-defined, antemedial fascia narrow, dilated at costa, hardly discernible but usually 
distinguished by slightly heavier fuscous suffusion; terminal fascia narrowly edged inwardly 
from costa to near tornus with pale buff ground colour, diffuse distally, containing 3 or 4 quadrate 
hair brown strigulae along costa interspersed with ground colour; cilia pale buff intermixed 
with a sprinkling of blackish. Hind wing light buff, obscurely mottled with narrow, sinuous, 
greyish lines ; cilia light buff, with a fuscous-black sub-basal line. 

Male genitalia (Fig. 63). Very similar to T. euglypta Meyrick but differing as follows: Transtilla 
with medial arm very short, well sclerotized and strongly denticulate at apex; aedeagus slightly 
longer than in euglypta but with apex not so strongly curved ventrad, containing two thorn-like 
cornuti as in euglypta. 

ruwenzori: Kimemba Camp, 11,900 ft., i.viii.1952 (Fletcher), J, holotype, genitalia slide 
3606; Lamia Valley, 11,900 ft., 30-31. vii. 1952 (Fletcher), 2 

The structure of the male genitalia of this species is very similar to that of euglypta, as mentioned 
above, and both species are superficially similar in general facies and are evidently closely related. 


Ruwenzori Expedition, 1952. Volume II, Number 12 

In hypsitropha the coloration is more sombre and the fore wing markings more obscure than in 
euglypta. Both species are remarkable for their huge size, and probably have the greatest wing 
expanse of the known species of this family. The only other member of the genus, T. chionochlaena 
Meyrick, which occurs in Abyssinia, is quite dissimilar superficially and in structure of male 
genitalia, and is considered as doubtfully congeneric. 


Capua pusillana (Walker) 

Tortrix pusillana Walker, 1863, Cat. Lep. Ins. B.M., 28 : 328. 
Capua exalbescens Meyrick, 1922, Zool. Meded. Leiden, 7:81. 
Epagoge probolias Meyrick, 1907, J. Bombay nat. Hist. Soc, 17 : 977. 
Capua metacentra Meyrick, 191 8, Exot. Microlepidopt., 2 : 163. 

Uganda: Semliki Forest, 2,850 ft. [Fletcher), 19 Bundibugyo, 3,440 ft. [Fletcher), 6 $9. 

Distribution: E. Africa (Uganda), S. Africa (Nyasaland), India (N. Bihar, Kumaon, Kashmir, 
Sikkim, Assam), Ceylon and Java. 

Capua liparochra Meyrick 

Capua liparochra Meyrick, 1928, Exot. Microlepidopt., 3 : 452. 

ruwenzori: Nawamba Valley, 6,500 ft. [Edwards), 1 3; Mahoma River, 6,700 ft. [Fletcher), 
6 3, 1 9; Nyamgasani Valley, 8,900 ft. (Buxton), 1 9. 

Previously known only from the type, a male collected in the Congo at 4,000 ft. The Ruwenzori 
specimens resemble the type in coloration and markings but are considerably larger, the male 
having a wing expanse of 20-22 mm., the female 24-25 mm., while the type measures 18 mm. 

The male genitalia and wing venation indicate generic affinities with Paramesiodes Diakonoff. 

Distribution: Congo. 

Capua spilonoma gitona subsp.n. (Figures 2, 64-66) 

Capua spilonoma Meyrick, 1932, Trans. R. ent. Soc. Lond., 87: 507. 

16-18 mm. General coloration of fore wing darker and more brownish-tawny than in nominate 
subspecies from Abyssinia, and wing expanse slightly less. Minor differences are evident in the 
male genitalia (Figs. 64-66) ; in gitona the keel-like projection a little distad of the middle of the 
sacculus is produced to a sharp thorn-like point, and in some examples the ventral edge of the 
sacculus tends to be irregularly serrulate; the lateral lobe (labis) of the transtilla is evenly rounded 
compared with the almost pyramidal outline in spilonoma. The solitary, prominent, acuminate 
lateral tooth on the apical section of the aedeagus in spilonoma is replaced in gitona by a low 
weakly serrulate ridge. 

ruwenzori: Mahoma River, 6,700 ft., 13-16.viii.1952 [Fletcher), 2 <$, including holotype, 
genitalia slide 8109; Misigo, 8,500 ft. [Fletcher), 2 


Parapandemis eustropha sp.n. (Figures 3, 67-69) 

2 24-25 mm. Labial palpus fermginom-brownish, suflused greyish above and below, paler 
inwardly; apex of terrninal segment yellowish. Head, patagia and tegula drab, patagia and tegula 
with ferruginous admixture. Thorax drab, darkened with strong admixture of blackish brown, 
a sprinkling of ferruginous scales anteriorly. Antennal scape warm buff, overlaid with purplish 
fuscous and ferruginous above except apical margin; flagellum light brown, clothed above widi 
warm buff scales, individual segments broadly banded with drab suffusion apically. Fore wing 
ground colour light buff, strongly overlaid with distinctive vinaceous drab, except for thin 
marginal lines bordering fasciae; markings well-defined outward-oblique nearly parallel fasciae, 
dark vinaceous drab partially overlaid with blackish-brown with weak ferruginous admixture ; 
basal patch outward-convex towards dorsum (inner margin) and reaching to middle, this and 
both margins of medial fascia and proximal margin of terminal fascia emphasized by dark 
ferruginous mixed with blackish; radial veins etched with blackish; cilia greyish mixed with 
ferruginous, a thin light buff basal line, a few cilia ripped with light buff. Hind wing light buff 
suffused greyish; cilia light buff, a dark grey sub-basal line, suflused warm buff between this and 
edge of wing, a broad somewhat diffuse greyish suffusion medially. 
Female genitalia (Figs. 67-69). 

rdwenzori : Mahoma River, 6,700 ft., 13-16.vhi.1952 (Fletcher), 2 2, hicluding holotype, 
genitalia shde 3532. 

A distinctive species having conspicuous well-defined tasciae on the fore whig reminiscent of 
P. isotetras (Meyrick), comb.n., from Fernando Po (W. Africa), to which it is closely related. 

Parapandemis orophila sp.n. (Figures 4, 70) 
5* 20 mm. Except for smaller size and shght differences in coloration this species fits the description 
of the previous species, P. eustropha Bradley, having a similar disposition of fore whig markings 
but with the outer (distal) margin of the basal patch or fascia almost straight and not convex and 
reaching to middle of inner margin (dorsum) as in eustropha. The ground colour of the fore wing 
is warm buff, extensively suffused with buffy brown mixed -with a trace of vinaceous drab - in 
eustropha the fore wing is extensively suffused with vinaceous drab, especially in the areas of the 
wring between the markings, and is very distinctive. Hind wing pale warm buff, diffuse greyish 
maculae distallv and around margins ; cilia light buff, suffused greyish apically. 
Male genitalia (Fig. 70.) 

kuwenzoei: Mt. Kinangop, 8,000 ft., xh.1934-i.1935 (Edwards), 1 3, holotype, genitalia 
shde 7048. 

Paramesiodes aprepta sp. n. (Figures 5, 7 I -75) 
S 17-23 mm., 2 19-24 mm Labial palpus approximately z\ times width of eye in male, 2 times in 
female; ochraceous-buff, mixed with fuscous exteriorly. Head, thorax and tegula warm buff or 
orchraceous-buff, sometimes suffused greyish. Antenna dark brown above, warm buff or light 
brown below, scape mixed with warm buff above; flagellum in male finely ciliate, ciliations 
approximately as long as width of shaft; female with ciliations vers 7 short; in bodi sexes a small 
patch of warm buff scales present at base of each segment dorsally from base to near apex; 111 the 
male these are roughened and more conspicuous. Fore wing general coloration and markings 


Ruwenzori Expedition, 1952. Volume II, Number 12 

variable: in male holotype and majority of paratypes as follows: ground colour greyish light buff, 
areas between markings lightly irrorate with brownish; markings brownish with a variable 
admixture of tawny, mostly diffuse and ill-defined ; basal marking broad, outer margin verv diffuse, 
strongly outward-convex at middle ; medial fascia stronger, outward-oblique, somewhat undulate, 
inner margin well-defined with coloration often darker and sometimes with an admixture of tawny 
and blackish producing a coffee coloured effect, outer margin mostly obliterate, traceable 
from costa, where fascia is very narrow, to end of cell area, strongly outward-oblique ; apical area 
of wing from about J on costa to termen a little above tornus suffused with greyish, slightly darker 
on costa, an indication of a broad pre-apical costal blotch, inner margin of apical area sometimes 
strengthened with a few specks of tawny or dark brown ; in female allotype and two female para- 
types the coloration is much darker and the general appearance of the fore wing is greyish fuscous 
with markings dark brown; in other specimens the markings are weak or obliterate; cilia con- 
colorous. Hind wing whitish or ochreous-white, sometimes with greyish suffusion, distal half 
weakly mottled with dark grey, most strongly towards apex; cilia concolorous. 

Male genitalia (Figs. 71, 72). Valva semimembranous, very short and broad, subovate, varying 
slighdy in outline, ventral margin normally straight as in Fig. 71, sometimes curved and valva 
more rounded distally as in Fig. 72. Sacculus narrow, slightly dilated at middle. Uncus narrowly 
spatulate. Socius well developed, membranous, finely haired, subquadrate. Transtilla well sclero- 
tized, moderately broad medially, dilated laterally and with latero-caudal margin produced and 
rounded, armed with bristly teeth decreasing in size towards comparatively narrow medial part 
of transtilla. 

Female genitalia (Figs. 73-75.) 

ruwenzori: Nyamaleju, 10,530 ft., 14-19.vii.1952 (Fletcher), 12 q, including holotype, geni- 
talia slide 7038; Nyinabitaba, 8.650 ft., 7-13.vii.1952 (Fletcher), 2 2, including allotype, genitalia 
slide 8144; Lamia Valley, 11,900 ft., 30-3i.vii.i952 (Fletcher), 1 3; Bigo, 11,400 ft., 20-22.vii.1952 
(Fletcher), 10 5*; Bahrungi River, 11,200 ft., i.vii.1952 (Fletcher), 1 Nyamgasani River, 
10,500 ft.-i 1,500 ft., i.1935 (Buxton), 1 3; Lake Mahoma, 9.600 ft., 12.vii.1952 (Fletcher), 2 o*5 
Mahoma River, 6,700 ft., 13-16.viii.1952 (Fletcher), 1 2. 

Larger than the two Malagasy species, P. longirostris Diakonoff and P. minor Diakonoff, the only 
other species previously known representing the genus. In those two species the uncus of the male 
genitalia is broad throughout its length and truncate distally, appearing rectangular in shape 
viewed ventrally; in aprepta the uncus is narrower basally and dilated and rounded distally. 

Niphothixa agelasta sp.n. (Figures 6, 76) 

o* 1 8 mm. Labial palpus fuscous exteriorly, whitish interiorly ; tips of scales on exterior lightened 
with pale drab-grey producing an irrorate effect, apex of terminal segment pale drab-grey. Head, 
thorax, tegula, patagia and antenna greyish fuscous, scales tipped with drab producing an 
irrorate effect. Fore wing ground colour warm buff, diffusely irrorate with greyish fuscous striae 
forming transverse chains especially in distal area; markings fuscous mixed with fuscous-black, 
basal patch diffuse, strongest at costa, outer margin obtusely outward-angulate before middle, 
weakly outlined with fuscous-black ; medial fascia outward-oblique from costa before middle to 
inner margin (dorsum) near tornus, containing a weak admixture of ochreous in central and 
dorsal areas, inner edge well-defined, sinuous and mixed with blackish, outer margin diffuse, not 
defined; a well-defined preapical marking on costa lightly sprinkled with ochreous and blackish; 

Microlepidoptera 87 

a few ochreous scales in striae between apex of preapical marking and tornus; cilia warm buff, a 
thin submedial drab-grey line along termen broadly edged with strong fuscous suffusion, a 
fuscous streak from inner margin before tornal angle. Hind wing drab, wbitisb along costal 
margin; cilia warm buff, a fuscous sub-basal line. 
Male genitalia (Fig. 76). 

stjwinzoei: Ibanda, 4,700 ft., 4-12.ix.1952 (Fletcher), 1 holotype, genitalia slide 7047. 

The genus Niphothixa Diakonoff was originally erected to accommodate two species from 
Madagascar. Of these, the species described above, N. agelasta, and also the following new species, 
are nearest N. niphadacra Diakonoff, having veins 3 and 4 of the hind wing connate. N. agelasta 
differs superficially from niphadacra in having the thorax a more or less uniform greyish fuscous, 
the apical fourth not being yellowish-white, and in the general coloration of the fore wing, 
which in niphadacra is a "deep fulvous-purple". 

Niphothixa ophina sp.n. (Figures 7, 77-78) 

o* 18-20 mm. Labial palpus basal and second segments snuff brown diffusedly irrorate with bister 
exteriorly, roughened dorsal scales suffused cinnamon-buff interiorly; terminal segment sayal 
brown diffusedly irrorate with bister. Head, thorax and tegula bister, a weak cinnamon-buff 
suffusion medially from crown of head to middle of thorax. Antenna fuscous, latero-posterior 
scaling bister basally becoming warm buff towards apex; scape light buff, overlaid with mummy 
brown above. Fore wing ground colour cinnamon, with extensive light vinaceous-drab suffusion; 
markings bister mixed with mummy brown ; basal marking occupying J, outer margin slightly 
outward-concave ; medial tascia from costa before middle, outward-oblique to inner margin 
(dorsum) before tornus, narrow and well-defined at costa, broadening and becoming obliterate 
beyond middle ; preapical triangular costal marking strong, emitting a diffuse bister streak from 
apex to termen parallel with medial fascia, a few scattered mummy brown striae distad and basad 
of this streak, a few thick short striae on inner margin between basal patch and medial fascia and 
along costa; radial veins finely etched with mummy brown penetrating into cilia as thickened 
dashes at tornus; an admixture of russet scales, not very conspicuous, along termen; cilia warm 
buff, a broad mummy brown medial band from apex to tornus, inner edge sharply denned, 
outer margin diffuse and extending into cilia. Hind wing pale warm buff, weakly infuscate in 
distal and vannal areas, some scattered stronger fuscous mottling at apex and along terminal 
margin; cilia concolorous, a broad wen-defined greyish brown sub-basal line, apices often whitish. 
At the base of the abdomen and most clearly visible in dissected specimens is a pair of subventral 
patches of specialised scales (Fig. 78). 

Male genitalia (Fig. 77). Sacculus broad, compressed, without tooth-like projection. Aedeagus 
long, curved throughout its length, 4 or 5 comparatively short blade-like cornuti. 

edwenzori: Mahoma River, 6,700 ft., 13-l6.viii.i952 (Fletcher), 1 & holotype, genitalia 
slide 3609; Misigo, 8,550 ft., 2-3 .vhi. 1952 (Fletcher), 1 $. 

Distinguished from the previous species by the dark coloration of the fore wing and the weaker 
contrast between the transverse fasciae and the ground coloration. 

Tortrix dinota Meyrick 
Tortrix dinota Meyrick, 191 8, Exot. Microlepidopt., 2 : 168. 
Uganda: Bundibugyo, 3,440 ft. (Fletcher), 1 q. 


Ruwenzori Expedition, 1952. Volume II, Number 12 

The male of this species is remarkable for the peculiar bifurcate aedeagus of the genitalia. This 
and certain other characters found in the male genitalia indicate that the species does not properly 
belong in the genus Tortrix L. (sensu stricto). 

Distribution: Nyasaland (bred from spun shoots of cotton, Gossypium). Kenya (bred from 
Arabian coffee, Coffea arabica), Ghana, Sierra Leone and Principe I. 

Tortrix edwardsi sp.n. (Figures 8, 79) 

cT 20 mm. Labial palpus warm buff, basal and second segments with a weak admixture of ochra- 
ceous-tawny exteriorly, terminal segment wholly suffused greyish. Head ochraceous-buft mixed 
with ochraceous-tawny. Thorax and tegula ochraceous-tawny, thorax strongly suffused hair 
brown, apical half of tegula suffused drab-grey. Antenna warm buff, scape suffused greyish above. 
Fore wing ochraceous-buff diffusedly marked with hair brown striae especially along costa, a 
moderately strong fuscous mixed with ferruginous fascia from costa before middle, narrow and 
moderately well-defined at costa, quickly broadening and forking to straddle a well-defined 
triangular cream colour marking, the base of which rests on inner margin (dorsum) from about 
j to a little beyond middle and with apex reaching to about middle of wing, distal furcation of 
fascia very broad and somewhat diffuse, outer edge slightly sinuate extending obliquely to dorsum 
near tornus, proximal furcation broadening gradually towards inner margin; triangular marking 
traversed vertically by several short striae similar in coloration to fascia; a poorly defined sub- 
triangular preapical marking resembling fascia, blended with a diffuse terminal fascia the inner 
margin of which is moderately well-defined and extends from apex of costal marking to termen 
near tornus and is parallel with outer edge of medial fascia; cilia (worn in type) concolorous. 
Hind wing cinereous; cilia cartridge buff, with a greyish sub-basal line. 

Male genitalia (Fig. 79). Uncus digitate, apex bluntly rounded, setose ventrally. Socius pendu- 
lous, hirsute. Gnathus a pair of compressed sclerotized arms with apices fused medially. Sacculus 
narrow, apex aciculate, a strong inward-projecting slender tooth-like projection near base, three 
slightly smaller tooth-like projections spaced equidistantly along inner edge of ventral margin. 
Transtilla labides globulose, bristling with strong denticulations, caudal margins connected 
medially by a short sclerotized rod-like bar. Aedeagus stout, orifice dorsal, apex bifurcate, upper 
prong produced to a long aciculate point, lower prong in the form of a strong tooth-like pro- 
jection opposite orifice, curved slightly ventrad. 

ruwenzori: Namwamba Valley, 6,500 ft., xii. 1934-1.1935 (Edwards), 1 <3\ holotype, 
genitalia slide 7195. 

Belonging to a species group which includes Tortrix meridionana (Walker), sp.rev., T. adustcma 
(Walsingham), sp.rev., and T. dorsiplagana (Walsingham), sp.rev., and superficially nearest 
the last species. These species are variable and may sometimes bear a close superficial resemblance 
to each other, so that edwardsi is more reliably distinguished by the strongly bifurcate apex of 
the aedeagus in the male. 

The three species mentioned above as allied to edwardsi are removed from synonymy, together 
with T. reciprocana (Walker), sp.rev., and T. capitana (Felder), sp.rev., under T. capensana 
(Walker), where they were placed by Meyrick (1908, Proc. zool. Soc. Lond., 1908 : 723). The 
re-examination of the types of capensana, meridionana, capitana and reciprocana, which are in the 
British Museum, has shown them to be distinct specifically; and examination of specimens of 
adnstana and dorsiplagana from Walsingham's collection (the types of the last two species not 

Microlepidoptera 89 

being available as they arc in the Museum at Cape Town) indicates that these arc also distinct 

Contrary to Meyrick's opinion (loc. cit), none of the above species is a true Tortri.x. T. capensana 
is generically distinct from the other species hitherto placed in synonymy with it and belongs in a 
genus near Capua. The other species, including cdwardsi, properly belong in a genus related to 
Ptycholonia Stephens and Leontochroma Walsmgham, but as they cannot be more accurately 
placed they are for the present retained in the genus Tortri.x. 

Tortrix chalicodes Meyrick 

Tortrix chalicodes Meyrick, 1920, Voyage de Ch. Alluaud et R. Jeanne] en Afrique, 2, Micro- 
lepidoptera, p. 48. 

ruwenzori: Misigo, 8,550 ft. {Fletcher), 1 $; Nyinabitaba, 8,650 ft. (Fletcher), 4 1 $; 
Nyamaleju, 10,530 ft. (Fletcher), 2 $\ Bigo, 11,400 ft. (Fletcher), 1 $. 

The additional Ruwenzori material shows chalicodes to be a true montane species, occurring 
at altitudes from a little under 6,000 ft. to over 1 1,000 it. At higher elevations the wing expanse 
increases considerably and the fore wing markings become obsolescent, the wings then being 
almost unicolorous. On average the female is larger than the male, the wing measurements for 
the Ruwenzori specimens being: q* 20-32 mm., $ 28-33 mm. 

This species does not belong in the genus Tortri.x, its generic affmites lying nearer Parapaudemis, 
but there appears at present to be no described genus to which it can be properly transferred. 

Distribution: Kenya. 

Tortrix stenophora sp.n. (Figures 9, 80-85) 

S $ 10-15 mm. Labial palpus ochraceous-buff, paler interiorly, second segment slightly infuscate 
exteriorly. Head, tegula and thorax ochraceous-buff, tegula lightly and thorax heavily suffused 
with drab. Antenna light buff, apices of segments spotted with blackish brown dorsaily; scape 
ochraceous-buff, infuscate dorsaily. Fore wing of male with moderate costal fold from base to 
near middle, ground colour in both sexes light buff diffusely irrorate with brownish and drab 
mixture, a few poorly defined scattered greyish strigulae and striae, darker along costa; markings 
yellowish light brown, suffused greyish; basal patch indicated by slight costal and dorsal suffusion; 
medial fascia outward-oblique from before middle of costa to dorsum near tornus, moderate, 
inner margin straight or slightly undulate, distal margin diffuse and dilated with drab suffusion 
in tornal proximity; preapical costal patch suffused greyish, produced inwardly as a rather narrow 
fascia extending obliquely towards termen, well defined along inner margin which is slightly 
undulate and edged with ochreous mixed with blackish and extends to tornus, diffuse and obscure 
distally and reaching termen before middle; cilia light buff or ochreous, a greyish sub-basal line. 
Hmd wing greyish drab; cilia concolorous or paler, a greyish sub-basal line. 

Male genitalia (Figs. 80-82). 

Female genitalia (Figs. 83-85). 

ruwenzori: Ibanda, 4,700 ft., 4-12.ix.1952 (Fletcher), 3 $, including holotypc, genitalia 
slide 7039, 2 including allotype, slide 8140. 

Closely related and superficially very similar to Tortrix scaeodoxa Meyrick. The male genitalia 
of both species are structurally similar but differ in the shape of the valva which is ovate 111 

90 Ruwenzori Expedition, 1952. Volume II, Number 12 

scaeodoxa and triangulate in stenophora, and in the uncus which in scaeodoxa is more elongate and 
produced distally to an acute point. Neither species strictly belongs in the genus Tortrix, and j 
the structure of the male genitalia indicates their generic affinities lie near Adoxophyes, but the 
separation of veins 7 and 8 in the fore wing, which in Adoxopliyes are stalked, excludes them from 1 
that genus. They are therefore provisionally retained in Tortrix, since no described genus can be I 
found which will properly accommodate them. 

Metamesia physetopa (Meyrick) 
Tortrix physetopa Meyrick, 1932, Trans, ent. Soc. Lond., 80 : 111. 
ruwenzori: Misigo, 8,500 ft. (Fletcher), 1 Nyinabitaba, 8,650 ft. (Fletcher), 1 
Distribution: Abyssinia. 

Metamesia elegans (Walsingham) 

Lozotaenia elegans Walsingham, 1881, Trans, ent. Soc. Lond., 1881 : 224, pi. 10, fig. 4. 
Cacoecia hedrastis Meyrick, 1908, Proc. zool. Soc. Lond., 1908 : 722. 
Cacoecia prona Meyrick, 1911, Ann. Transv. Mus., 2 : 223. Syn.n. 

ruwenzori: Ibanda, 4,700 ft. (Fletcher), 3 q\ 

The above synonymy has been established following the comparison of type material of 
M. hedrastis (Meyrick) and M. prona (Meyrick), and topotypical specimens of M. elegans deter- 
mined by Walsingham. All three species were described from South Africa. The type of elegans \ 
is in the South African Museum at Cape Town, and has not been examined during the present 
study. The type of prona is in the British Museum (lectotype designated by Clarke, 1958, Catalogue 
of the Type Specimens of Microlepidoptera in the British Museum (Natural History) described 
by Edward Meyrick, 3 : 252, pi. 126, figs. i-ib). 

Cacoecia hedrastis Meyrick was synonymised under Tortrix elegans by Meyrick (1912, Lep. Cat., 
10 : 33), but neither a type nor subsequent lectotype designation appears as yet to have been 
made for this species. A male in the Meyrick collection in the British Museum, representing one I 
of the two original syntypes, is therefore now selected as LECTOTYPE. It bears labels with the i 
following data, "Pretoria, Transvaal, Janse 3.06" "E. Meyrick coll." "Genitalia slide No. 8054". 

Distribution: M. elegans occurs in S. and E. Africa; examples are known from Natal, Pretoria, | 
Transvaal, Mashonaland, Kenya, Uganda and the Congo. 

Metamesia octogona sp.n. (Figures 10, 86) 

o* 16-19 mm. Labial palpus light buff interiorly except terminal segment; clove brown mixed 
with ochraceous-buff exteriorly including whole of terminal segment. Head greyish hair brown, 
crown and chaetosema suffused with ochraceous-buff. Antenna light buff below, warm buff 
above, scales roughened postero-dorsally, tipped with fuscous or black; scape light buff below, 
warm buff above, marked with mummy brown dorsally except at apex. Thorax and tegula olive 
brown with ochreous admixture, tegula slightly paler towards apex. Fore wing with strong 
costal fold to near middle; ground colour light drab, a few dispersed olive brown dots or striae 
edged and mixed with raw sienna; markings basically olive brown, as follows: a moderate 
diffuse basal patch mixed with olive brown extending into costal fold; medial fascia well defined, 
narrow at middle of costa quickly dilating and resting on inner margin (dorsum) from about f 

Microlepidoptera QI 

to tornus, almost completely suffused with vinaceous-grey mixed with ochrcous except at 
margins, inner (proximal) margin straight, slightly outward-oblique from costa, outer (distal) 
margin curved outwards from below costa, continuing in a bold curve to tornus, a nearly 
constant admixture of raw sienna distributed along both margins and forming a weak yellowish 
demarcation line; a similar but somewhat broken and indefinite pre-apical marking on costa, 
followed by an obliquely transverse chain of dark olive-brown striae from near apex of marking 
to termen above tornus, similar scattered striae in terminal area beyond, surrounded by vinaceous- 
grey suffusion overlaying almost entire terminal area; cilia ochraceous-buff, a distinct broad 
vinaceous-grey sub-basal line, broadening and becoming diffuse around tornus. Hind wine; drab, 
whitish along costa towards base; cilia light buff, with a strong, drab sub-basal line. 

Male genitalia (Fig. 86). The caudal margin of VHIth sternite bears an array of eight very 
large, curved, prong-like setae, arranged four at each side. In the specimen shown in Fig. 86, 
three of the left-hand group of setae are missing. 

ruwenzori: Nyinabitaba, 8,650 ft., 7-13.vii.1952 (Fletcher), 1 Q A , holotype, genitalia slide 
4507; Mahoma River, 6,700 ft., 1 3-16. vii. 1952 (Fletcher), 1 £. 

The structure of the male genitalia of this species indicates close relationship with Metatnesia 
endopyrrha (Meynck), comb.n., in which the Vllftb sternite is similarly developed and bears a 
similar number of prong-like setae. The two species differ superficially in coloration of the hind 
wing, octogona lacking the orange coloration present in endopyrrha. 

Procrica ophiograpta (Meyrick) 
Tortrix ophiograpta Meyrick, 1932, Trans, ent. Soc. Lond., 80 : no. 
ruwenzori: Ibanda, 4,700 ft. (Fletcher), 1 6*. 
Distribution: Abyssinia. 

Epichoristodes panochra sp.n. (Figures n, 87-88) 

o* 14-16 mm. Labial palpus whitish suffused exteriorly with fuscous. Head, thorax and tegula 
whitish suffused with greyish fuscous. Antenna clothed with whitish scales dorsally; scape wholly 
whitish. Fore wing ground colour light buff, lightly suffused with greyish fuscous, suffusion 
stronger in basal area especially along costal margin; distal third of wing suffused with fuscous 
forming a broad somewhat diffuse fascia the inner edge of which is obtusely angled towards 
cell; cilia whitish. Hind wing whitish tinged with greyish; cilia whitish. 

Male genitalia (Figs. 87-88). Lateral shoulder of transtilla crowned with slender crenellate 
column. Aedeagus without cornuti 111 specimens examined but slight internal scars indicate 
deciduous cornuti; apex obliquely tapered, rim of orifice minutely denticulate. 

ruwenzori: Misigo, 8,550 ft., 2-3. vii. 1952 (Fletcher), 2 6\ including holotype, genitalia 
slide 7514. 

Similar to Epichoristodes lianaea (Meyrick), comb.n., but distinguished by the diffuse trans- 
verse fascia in distal area of fore wing. 

92 Ruwenzori Expedition, 1932. Volume II, Number 12 

Epichoristodes atycta sp.n. (Figures 12, 89-90) 

6* 14-15 mm. Labial palpus ochraceous-buff, second segment suffused with fuscous exteriorly 
except at apex. Head warm buff or ochraceous-bufF, thorax and tegula ochraceous-bufF, mixed 
with brownish anteriorly. Antenna warm bufF, segments dotted with dark brown dorsally. Fore 
wing warm bufF with irregular diffuse Fuscous suffusion; markings Fuscous, difFuse and obscure; 
an indication oF a basal patch strongest along costal margin; medial Fascia hardly discernible, 
outward-oblique From about middle oF costa, outer margin connected by Fuscous suffusion to a 
slightly stronger preapical patch on costa; medial veins partially outlined with fuscous intermixed ! 
with a few scattered blackish specks; a small well-defined blackish dot in upper part of cell at J, 
preceded obliquely below by a similar weaker dot in plical fold, and followed by another slightly 
larger dot in plical fold a little above dorsum at about § , a fourth incomplete dot on inner margin 
at about f ; cilia warm buff, paler at apices. Hmd wing light buff, basal half overlaid with pro- 
nounced dark fuscous suffusion except along costal margin; cilia light buff. 

Male genitalia (Figs. 89-90). Very similar to those of E. panochra Bradley, but differing as 
follows: denticulate cluster on dilated lateral portion of transtilla with teeth longer and base 
shallower; aedeagus similarly tapered towards apex but with rim of tapered portion smooth 
and not minutely denticulate as in panochra, a dense internal sheaf of hair-like cornuti. 

ruwenzori: Mahoma River, 6,700 ft., 13-16.viii.1952 (Fletcher), 3 6*, including holotype, 
genitalia slide 5206. 

Near E. panochra, from which it may be distinguished superficially by the dark fuscous suffusion 
in the basal half of the hind wing. 

Epichoristodes heterotropha sp.n. (Figures 13, 91-92) 

18 mm. Labial palpus warm buff, second segment diffusedly irrorate with greyish, sprinkled 
with ferruginous exteriorly; terminal segment suffused greyish, an admixture of ferruginous at 
apex. Head, thorax and tegula warm buff, scales tipped with pale vinaceous-fawn especially on 
tegula. Antenna warm buff weakly suffused with fuscous towards base posteriorly. Fore wing 
warm buff lightly strigulatc with brownish; markings somewhat diffuse, tawny; basal patch | 
very broad, reaching to \, strong in costal half and darkened with blackish strigulae, very weak ; 
and difFuse towards inner margin (dorsum) ; medial Fascia oblique From middle oF costa to inner 
margin near tornus, narrow and strongest at costa and darkened with blackish, dilate in discal I 
and tornal areas; a subtriangular costal patch at I with a weak blackish admixture, emitting a 
stria From its apex to tornus; cilia warm bufF at apices, tawny basally, a difFuse greyish medial 
line. Hind wing pale buff-yellow, costal margin broadly white to near apex, basal third oF this 
whitish area irrorate with purplish black scales; cilia cartridge bufF, a basal line concolorous 
with wing, bordered outwardly by a slightly darker thin sub-basal line, greyish around apex, 
sub-basal line tawny at apex. General coloration of underside of fore wing ochraceous-buff and 1 
hind wing pale warm buff, basal area of costa of both wings with an extensive patch of sparsely 
distributed purplish black scales, sometimes partly concealed by translucent whitish scales; a 
sprinkling of similar blackish scales either side of median vein towards base of fore wing; vannal 
margin of hind wing narrowly edged with similar black scales above and below. Legs whitish, 
fore and middle legs suffused with ochraceous-buff exteriorly, tibia and tarsal segments of fore 
leg marked with diffuse greyish fuscous bands. A well developed purplish brown hair-tuft 
arising from thorax laterally immediately below wings. 

Microlepidoptera 93 

Male genitalia (Figs. 91, 92). Uncus elongate-spatulate, bristled on ventral surface of spatulate 
portion. Gnathus arms not fused medially. Socius large, pendulous. Transtilla very narrow at 
middle. Labis greatly enlarged, simple, sparsely setose antcro-ventrally. Aedeagus curved vcntrad, 
apex oblique, produced ventrally to a slender, tapered lip; a dense, compact sheaf of numerous 

ruwenzori: Ibanda, 4,700 ft., 20-21. vhi. 1952 (Fletcher), 1 <J, holotype, genitalia slide 4506. 

This species may tentatively be placed next to E. atycta. The separation of the gnathus arms, 
which are free and not fused medially as in the male genitalia of atycta and other related species, 
is atypical of the genus. 

Acleris thylacitis (Meyrick) 

Peronca thylacitis Meyrick, 1920, Voyage de Ch. Alluaud et R. Jeannel en Afnque Orientale, 
2, Microlepidoptera, p. 57. 

ruwenzori: Mahoma River, 6,700 ft. (Fletcher), 1 Nyamaleju, 10,530 ft. (Fletcher), 2 r j\ 
3 $; Bigo, 11,400 ft. (Fletcher), 1 6*. 

A montane species exhibiting considerable variation. The Ruwenzori specimens include forms 
similar to those originally described by Meyrick, including a male and female of the form having 
a semioval blotch on the apical f of the costa of the fore wing. 

The type (a lectotype female selected by Dr. P. Viette) in the Paris Museum has been compared 
with a female from Nyamaleju (genitalia slide 3436), which it resembled most closely super- 
- ficially, and found to be identical in genitalia. 

Distribution: E. Africa. 

Crocidosema plebejana Zeller 
Crocidosema plebejana Zeller, 1847, Isis von Oken, 40 : 721. 
Uganda : Semliki Forest, 2,850 ft. (Fletcher), 1 $. 

Distribution: Cosmopolitan in tropical and subtropical regions, and sometimes occurring in 
temperate zones. Larva on seeds of Malvaceae. 

Eucosma phaeochyta sp.n. (Figures 14, 93) 

cJ 22-23 nim. Labial palpus grey, second segment paler interiorly, infuscate exteriorly, brush 
partly concealed and reaching beyond apex of terminal segment which is also infuscate except 
at apex; sometimes a trace of pale ferruginous in upper margin of second segment. Head, thorax 
and tegula grey, sometimes with a ferruginous admixture, front of head usually paler. Antenna 
dark grey, minutely ciliate. Fore wing pale greyish ochreous, overlaid with irregular scattered 
brown markings, heaviest distally, diffuse and forming no definable pattern; a variable admixture 
: of scattered pale ferruginous scales, most evident in distal area, along costa and termcn; ocellar 
area above tornus sometimes free from extraneous coloration; an elongate diffuse leaden-grey 
patch from costa before apex to upper margin of cell, usually with brownish coloration super- 
imposed; four or five jet black post ocellar dots, forming quadrangle; cilia greyish, sometimes 
brownish below apex to near middle of termcn, an admixture of ferruginous along termcn to 
near tornus. Hind wing with veins 3 and 4 connate; light grey; cilia concolorous, with darker 
grey sub-basal shade. 
Male genitalia (Fig. 93). 

94 Ruwenzori Expedition, 1952. Volume II, Number 12 

ruwenzori: Nyamaleju, 10,530 ft., 14-19.vii.1952 [Fletcher), 2 including holotype, 
genitalia slide 6542; Lake Mahoma, 9,600 ft., 12.vii.1952 (Fletcher), 1 

Near E. orthopeda Meyrick, but having appreciably greater wing span, orthopeda measuring 
only 18 mm., and further distinguished by the overall darker coloration of the fore wing and 
the obscure wing pattern. 

Notocelia scotodes sp.n. (Figures 15, 94) 

o* 16 mm. Labial palpus white, basal and second segments overlaid with fuscous-black exteriorly 
and beneath. Head with lower part of face white, narrowly bordered with fuscous around margin 
of eye; crown and vertex fuscous-black, individual scales tipped with greyish white. Thorax and 
tegula fuscous-black, middle of thorax and apical half of tegula mixed with whitish. Fore wing 
white, costa irregularly strigulated dark fuscous, a weak sprinkling of ochreous scales in and 
below costal margin; markings fuscous mixed with fuscous-black, a small somewhat diffuse 
basal patch, margin angled outwards at middle; a broad antemedial fascia at j, outer edge sharply 
defined, irregular, slightly outward-oblique, inner edge diffuse; a broad, diffuse, ill-defined 
fuscous-black mixed with leaden-grey postmedial fascia, preceded by an irregular fuscous-black 
line from costa to inner margin (dorsum) ; a circular fuscous-black apical marking below which 
extends a broad fuscous-black mixed with grey suffusion along termen; white discal area with 
irrorate leaden-grey suffusion, heavier towards inner margin; cilia dark grey, a whitish basal 
line, apices of some cilia whitish grey. Hind wing fuscous; cilia grey, a thin whitish basal line 
followed by broad sub-basal shading. 
Male genitalia (Fig. 94). 

ruwenzori: Mahoma River, 6,700 ft., 13-16. vii.1952 (Fletcher), 1 3, holotype, genitalia 
slide 5199. 

Apparently the only representative belonging to the genus Notocelia known from Uganda. 
Readily distinguished from N. suffusana Duponchel, the geographical range of which extends 
southwards to North Africa, by the appreciably darker coloration of the fore wing. 

Crimnologa fletcheri sp.n. (Figures 16, 95-97) 

cj $ 26-33 mm. Labial palpus with terminal segment wholly white, second segment black excepting 
white apical half of dorsal margin, basal segment black above, white below. Head wholly pale 
sulphur yellow. Antenna black, scape white below, flagellum with a thin white line ventrally 
from base to about §. Thorax and tegula pale sulphur yellow, a heavy quadrate black marking 
occupying most of anterior half of thorax, narrow anteriorly, broadening to full width of thorax 
at middle ; a prominent patch of raised black scales posteriorly on thorax ; basal half of tegula 
overlaid with black. Fore wing pale sulphur yellow (in worn specimens this colour fades and 
becomes whitish), an intricate broken pattern of small black markings as illustrated in Fig. 16; 
these markings vary slightly, in some examples certain of the costal markings may be fused and 
the apex of the triangular marking at middle of termen and the inner margin of the costal marking 
opposite it may be connected and form an oblique fascia; cilia pale sulphur yellow mixed with 
whitish at apex and tornus, wholly black along termen, on inner margin before tornus and on 

Microlepidoptera 95 

costa before apex. Hind wing grey, suffused brownish distally ; cilia brownish grey, whitish apic- 
ally, a dark brown sub-basal line. 

Male genitalia (Fig. 95). 

Female genitalia (Figs. 96, 97). 

ruwenzori: Nyamaleju, 10,350 ft., 14-19. iv. 1952 (Fletcher), 1 J, holotype, genitalia slide 
3239, 3 ?, including allotype, genitalia slide 3433, Bigo, 11,400 ft., 20-22. vii. 1952 (Fletcher), 1 
Namwamba Valley, 10,200 ft., xii.1934-i.1935 (Edwards), 1 

The only other known species belonging to this genus is C. perspicua Meyrick, described from 
a unique female collected on Kilimanjaro. The type has not been available for examination 
during the present study; from the description it resembles fietcheri in size and general appearance 
but differs in the pattern of the black markings of the fore wing, notably in the more numerous 
markings on the costa which has about eighteen transverse marks or small dots. 

Endothenia alpigena sp.n. (Figures 17-19, 98-102) 

o* $ 15-20 mm. Labial palpus fuscous-black, second segment mixed above and below (except 
apical brush) with yellowish white varying to greyish yellow (amber). Head fuscous-black mixed, 
sometimes very strongly, with light yellow or greyish yellow. Thorax fuscous-black; tegula 
fuscous, sometimes with an admixture of yellowish, normally with apex strongly suffused with 
greyish yellow varying less frequently to yellowish white. Antenna fuscous-black above, yellow- 
ish white beneath; in male finely ciliate. Abdomen fuscous or greyish fuscous above, caudal 
margins of segments fringed with long yellowish white scales ; underside yellowish white ; anal 
tuft of male yellowish white. Fore wing variable in pattern : fuscous mixed with fuscous-black, 
sparsely sprinkled with ferruginous-orange scales, more heavily in some examples (particularly 
allotype) than in others, an irregular whitish yellow fascia at \, narrow and angled outwards at 
middle, as in holotype (Fig. 17), or broad and diffuse as in allotype (Fig. 18); a second more 
extensive fascia from costa at §, constricted at middle thence divided, one fork reaching termen 
immediately above tornus, the other fork going to inner margin (dorsum) a little before tornus, 
in some examples (Fig. 19) these fasciae are partially or entirely obliterated by fuscous suffusion; 
a small almost invariably well-defined discal spot is present at end of cell, in some examples this 
is bordered costally with a few additional ferruginous-orange scales; costa irregularly marked 
with fuscous-black strigulae; small white interneural or neural dots spaced along termen which 
may sometimes have a heavier concentration of ferruginous-orange ; cilia fuscous, a strong fuscous- 
black basal band from apex to tornus inclusive, cilia along termen tipped with yellowish-white 
or greyish yellow. Hind wing white, apical area dark fuscous, in some examples the normally 
clear white area of the wing may be suffused with greyish; cilia white, a broad dark fuscous 
sub-basal line from above apex to below middle of termen; cilia along this line suffused with 
fuscous to near apices. 

Male genitalia (Figs. 98, 99). 

Female genitalia (Figs. 100-102). 

ruwenzori: Nyamaleju, 10,530 ft., 14-19.vh.1952 (Fletcher), 18 <J, including holotype, 
genitalia slide 10488, 7 $, including allotype, genitalia slide 10530; Lake Bujuku, 13,050 ft., 
22-28.vii.1952 (Fletcher), 4 <?, 1 $ (all with white fasciae on fore wing obliterate) ; Lake Mahoma, 
9,600 ft., 12.vii.1952 (Fletcher), 1 $; Stuhlmann Pass, 13,500 ft., 27.vi1.1952 (Fletcher), 2 $ (white 


96 Ruwenzori Expedition, 1952. Volume II, Number 12 

markings obliterate in both specimens); Mt. Stanley, 14,500 ft., 26.vii.1952 [Fletcher), 2 1 $ 
(white markings obliterate). 

Near E. vasculigera Meyrick, from which it may be distinguished by the white hind wing, 
which in vasculigera is dark grey. 

Epinotia penthrana sp.n. (Figures 20, 103-106) 

o* $ 29-34 mm. Labial palpus greyish sepia, second segment strongly suffused with golden yellow 
interiorly. Head greyish sepia, slightly darker on frons; sometimes an admixture of golden yellow, 
apices of individual scales on crown often pale grey. Antenna sepia, scape somewhat darker. 
Thorax and tcgula blackish sepia mixed with ferruginous, sometimes a sprinkling of white- or 
grey-tipped scales on thorax, especially in posterior raised scale-tuft; apex of tegula whitish or 
greyish. General coloration of fore wing blackish sepia, irregularly mixed with ferruginous, 
sometimes sufficiently strong to form streaks and patches, but in other examples partly obscured 
by bluish grey suffusion; coloration of markings varying from whitish to dark brown: numerous 
small strigulae, usually in pairs, scattered along costa; a moderately broad transverse fascia from 
costa at ^ to dorsum beyond middle containing a variable mixture of dark fuscous and ferru- 
ginous; a similarly coloured marginal band from immediately below apex, inner edge slightly 
sinuous to tornus, thence deflected inwards and becoming somewhat obscure and terminating at 
medial fascia a little above inner margin (dorsum) ; a small discal dot at end of cell; cilia matching 
general coloration of fore wing : greyish sepia with dark sub-basal line, extreme tips greyish or 
brownish. Hind wing whitish grey varying to grey, darker distally and sometimes with diffuse 
dark grey patches in terminal margin; cilia whitish grey, a greyish sub-basal line, a similar more 
diffuse subapical line. 

Male genitalia (Fig. 103). 

Female genitalia (Figs. 104-106). 

ruwenzori: Nyamaleju, 10,530 ft., 14-19.viii.1952 (Fletcher), 2 including holotype, 
genitalia slide 3229, 1 §, allotype, slide 3528; Bigo, 11,400 ft., 20-22.vii.1952 (Fletcher), 2 

Superficially nearest E. rhodochrauta (Meyrick) andf?. uimbosa (Meyrick), but at once distinguished 
from these and other known species in this group by its large size. 

Olethrentes phyllodoxa (Meyrick) 

Argyroploce phyllodoxa Meyrick, 1932, Exot. Microlepidopt., 4 : 310. 

ruwenzori: Misigo, 8,550 ft. (Fletcher), 1 $; Nyinabitaba, 8,650 ft. (Fletcher), 2 1 
Previously known only from the type. 

Distribution: Uganda (Ruwenzori). 

Olethreutes orestera sp.n. (Figures 21, 22, 107-110) 

S 24-27 mm., $ semibrachypterous, 18-20 mm. Length of labial palpus approximately 3 times 
width of eye, second segment extremely long, terminal segment very short and partly concealed 
by apical brush of second segment; general coloration medium grey, diffusedly irrorate with 
fuscous exteriorly and beneath; projecting bristly scales at apex of second segment dark fuscous; 
scales along dorsal margin of all segments tipped with whitish. Head, thorax and tegula medium 
grey; scales on head tipped with whitish; thorax suffused with dark fuscous posteriorly; tegula 

Microlepidoptera 97 

strongly suffused with dark fuscous. Antenna minutely ciliate in both sexes, greyish fuscous, 
apices of segments fringed with roughened scales, dark fuscous above with trace of purplish sheen, 
light grey below; scape dark fuscous. Fore wing in male subtriangular, apex prominent and in 
some examples examined showing strong tendency to turn upwards, termcn sinuate, subcostal 
and discoccllular veins separate, a circular cavity containing specialized scales situated on underside 
of wing and enclosed in basal furcation of vein ia; ground colour medium grey, sometimes 
weakly purplish, diffusedly irrorate with fuscous and fuscous-black; a blackish suffusion at base 
of costa, a broad longitudinal diffuse blackish band along plical fold from base to end of cell, 
paralleled by a narrower band along subcostal vein, both sometimes interrupted near middle, or 
in heavily marked examples dilate and merging medially so as to form an almost solid blackish 
central area, in any event both bands dilate and coalesce distally, thence turning obliquely costad 
and continuing as a narrow gradually tapering streak to apex, upper margin of distal half of 
subcostal band and whole of apical streak bordered with white, sometimes a sprinkling of ferru- 
ginous scales in blackish streaks, especially along edges; a sprinkling of whitish scales along termen; 
cilia grey, tipped with whitish; a diffuse dark greyish fuscous basal line. Fore wing in female 
greatly reduced, apex attenuate; general coloration similar to male but with markings mostly 
obsolete. Hmd wing in both sexes whitish grey; cilia concolorous, weak greyish basal and 
apical lines. 

Male genitalia (Fig. 107). 

Female genitalia (Figs. 108-110). 

ruwenzori: Lake Bujuku, 13,050 ft., 22-28. vii. 1952 (Fletcher), 2 $, including holotype, 1 $, 
allotype, genitalia slide 9610; Bigo, 11,400 ft., 20-22. vii. 1952 (Fletcher), 5 Stuhlmann Pass, 
13,500 ft., 27.vii.1952 (Fletcher), 1 <J, I $; Lamia Valley, 11,900 ft., 30-31. vii. 1952 (Fletcher), 2 
Kimemba Camp, 11,900 ft., i.vh.1952 (Fletcher), 1 J; Nyamaleju, 10,530 ft., 14-19.vii.1952 
(Fletcher), 1 S- 

The longitudinal markings of the fore wing, well-defined in the male and obsolescent in the 
female, and the reduced development of the fore wing in the female, make orestera a species of 
distinctive and unusual appearance. It is similar in size to O. phyllodoxa (Meyrick) with which it is 
provisionally associated until such time as it can be more precisely placed generically. Apart from 
similarity in size, the two species are quite distinctive and readily separated by difference in general 
coloration of the fore wing, orestera lacking the greenish coloration prevalent in phyllodoxa. 


Psamathocrita doloma sp.n. (Figures 23, 24, 111-114) 
cT $ 7-9 mm. Labial palpus yellowish white, basal and second segments diffusely irrorate with 
sepia, most strongly exteriorly, except at apex of second segment, which is encircled by a com- 
paratively whitish amiulus; terminal segment with broad basal and postmedial sepia annuh. 
Head, thorax and tegula yellowish brown with variable, sometimes very strong, greyish ad- 
mixture, front of head (face) usually much paler. Antenna pale ochreous basally, scape lightly 
suffused with greyish sepia above, flagellum suffused with greyish above basally and with 
alternate segments blackish to near middle, from about middle to apex the markings on the 
iged so that three adjacent blackish segments arc followed by three white 

9 8 

Ruwenzori Expedition, 1952. Volume II, Number 12 

segments interspersed with two blackish segments followed by three consecutive black segments 
followed by one white; the same sequence, beginning with three blackish segments is then re- 
peated to apex. Fore wing yellowish brown suffused with dark greyish; markings diffuse, obscure, 
consisting of blackish sepia liberally mixed with plumbeous; a small black dot of raised scales 
above tornus confluent with a smaller whitish patch basad; cilia grey. Hind wing and cilia grey. 

Male genitalia (Fig. 111). 

Female genitalia (Figs. 11 2- 11 4). 

ruwenzori: Mahoma River, 6,700 ft., 13-16.viii.1952 (Fletcher), 15 including holotype, 
genitalia slide 10148, 6$, including allotype, genitalia slide 10144. 

Uganda: Semliki Forest, 2,850 ft., 22.viii.-3.ix. 1952 (Fletcher), 1 1 <j>. 

Tentatively assigned to the Palaearctic genus Psamathocrita on account of the similarity of the 
genitalia structure, but superficially very different from other species in the genus, which are 
comparatively plain and whitish in appearance. 

Aristotelia epacria sp.n. (Figures 25, 115-119) 

6* $ 16-19 mm. Labial palpus white, second segment suffused fuscous to near apex, except dorsal 
margin and narrow interruption at middle; terminal segment with broad fuscous-black subapical 
and sub-basal annuli. Head whitish, diffusedly irrorate greyish fuscous. Antenna greyish fuscous, 
alternate segments suffused fuscous-black; scape suffused fuscous-black. Thorax and tegula 
whitish, suffused greyish fuscous, darker and approaching fuscous-black on thorax medially and 
towards basal half of tegula. Fore wing white, variably suffused, in places strongly, with greyish 
fuscous, markings blackish; a small blackish patch at base of costa, a narrow outward-oblique 
somewhat diffuse blackish fascia from costa at | terminating before inner margin (dorsum), 
followed distally by whitish area very weakly suffused with fuscous; a wedge-shaped blackish 
patch on costa at about J, contiguous with small spot of somewhat roughened black scales at 
about J across wing, a similar spot diagonally beyond this near middle of wing, a little below 
and slightly basad of which is an elongate blackish stigma lying in plical fold; a small spot of 
roughened blackish scales at end of cell a little above tornus; area between this and blackish spots 
at middle predominantly whitish; apical area mostly whitish, weakly suffused with greyish 
fuscous; scattered blackish scales along termen, around apex and in apical margin before apex; 
cilia greyish. Hind wing greyish; cilia matching or paler. 

Male genitalia (Figs. 115, 116). 

Female genitaha (Figs. 11 7- 11 9). 

ruwenzori: Lake Mahoma, 9,600 ft., 12.vii.1952 (Fletcher), 1 holotype, genitalia slide 
8717, 2 $, including allotype, genitalia slide 9616; Bigo, 11,400 ft., 20-22.vii.1952 (Fletcher), 1 6*; 
Lamia Valley, 11,900 ft., 30-31.vii.1952 (Fletcher), 1 1 $; Balirungi River, 11,200 ft., i.viii.1952 
(Fletcher), 1 cT- 

Tentatively placed near A achyrobathra Meyrick, from which it may be distinguished by slightly 
larger size and absence of ochreous coloration in the fore wing. 

Tricyanaula metallica Walsingham 
Tricyanaula metallica Walsingham, 1891, Trans, ent. Soc. Lond., 1891: 97, pi. 4, f. 34. 
congo: Gorges de la Pelenge, 1,150 m., (G. F. de Witte), 34 <$ ?• 
Distribution: Gambia, Transvaal and Madagascar. 



Chelaria stasimodes Meyrick 
Chelaria stasimodes Meyrick, 193 1, Exot. Microlepidopt., 4 : 70. 
Uganda: Bundibugyo, 3,440 ft. (Fletcher), 1 <J. 
Distribution: Mozambique. 


Timyra floccula sp.n. (Figures 26, 120, 121) 

eJ 22 rrrrn. Labial palpus buff yellow, second segment with very long erectly expanded hairs 
above. Head, thorax and tegula drab. Antennal flagellum buff yellow, a deep notch at base 
anteriorly; scape buff yellow, elongate, projecting scales at apex posteriorly. Fore wing buff 
yellow, anterior edge of costa slightly deeper yellow; markings ochreous, very diffuse and 
poorly defined; basal area of wing to near middle lightly suffused with ochreous, suffusion 
extending along costal margin towards apex and spreading into distal area; the semblance of a 
weak ill-defmed brownish fascia from below costa before middle, slightly outward-oblique to 
inner margin (dorsum) ; a similar indistinct fascia from below costa at f to inner margin before 
tornus ; cilia ochreous. Hind whig maize yellow, an admixture of buff yellow in distal area and 
below median vein to base, an elongate area of greyish brown scales occupying cell at middle of 
wing ; cilia pale maize yellow. 
Male genitalia (Figs. 120, 121). 

ruwenzori: Nyinabitaba, 8,650 ft., 7-13. vii. 1952 (Fletcher), 1 holotype, genitalia slide 

Related to T. aeolocoma Meyrick, which occurs in Malawi. The two species aresimilarin general 
coloration, but floccula may be readily distinguished by the absence of brown in the cilia of the 
fore wing, and the absence of the conspicuous greyish brown triangular apical patch, present on 
the hind wing in aeolocoma. 


Cosmopterix transcissa Meyrick 
Cosmopterix transcissa Meyrick, 1914, Exot. Microlepidopt., 1 : 203. 

Uganda : Semliki Forest, 2,850 ft. (Fletcher), 4 ex.; Bundibugyo, 3,440 ft. (Fletcher), 1 ex. 

congo: Lusinga, 1,700 m., 15.iv.1947 (G. F. Ae Witte), 4 ex.; R. Mubale, 1,480 m., 6.V.1947 
(G. F. de Witte), 1 <J. 

Distribution: Malawi (Mt. Mlanje). 

Labdia oxychlora Meyrick 
Stagmatophora promacha (Meyrick); Lefroy, 1909, Indian Insect Life, p. 536. Calcutta. 
Stagmatophora promacha (Meyrick) ; Meyrick, 1914, Ent. Mitt.-Suppl. no. 3 : 54- 
Pyroderces promacha Meyrick; Fletcher, 1920, Rep. Proc. ent. Meet. Pusa, i: 150. 

ioo Ruwenzori Expedition, 1952. Volume II, Number 12 

Pyroderces promacha Meyrick; Fletcher, 1921, Mem. Dep. Agric. India ent. Ser., vi: 100. 

Labdia promacha (Meyrick); Meyrick, 1927, Insects, Samoa, iii: 90. 

Labdia promacha (Meyrick); de Joannis, 193 1, Annls. Soc. ent. Fr., 98 : 728. 

Labdia oxycidora Meyrick, 1932, Exot. Microlepidopt., 4 : 314. 

Labdia promacha (Meyrick); Fletcher, 1933, Scient. Monogr. Counc. agric. Res. India., no. 4 : 3. 
Labdia fietchcreUa Bradley, 195 1, Ann. Mag. nat. Hist., (12) 4 : 510. 

Uganda: Bundibugyo, 3,440 ft. {Fletcher), 7 J 1 ?; Semliki Forest, 2,850 ft. [Fletcher), 8 $ $. 

Distribution: W. Africa, India and Ceylon. 

Pyroderces embrochota Meyrick 
Pyroderces cmbrocliota Meyrick, 1914, Exot. Microlepidopt., 1 : 280. 
Uganda: Bundibugyo, 3,440 ft. (Fletcher), 1 
Distribution: Sierra Leone. 

Pyroderces orphnographa Meyrick 
Pyroderces orphnographa Meyrick, 1936, Exot. Microlepidopt., 5 : 29. 
Uganda: Bundibugyo, 3,440 ft. (Fletcher), 1 
Distribution: Congo. 

Limnaecia pamphaea sp.n. (Figures 27, 122, 123) 

15-16 mm. Labial palpus yellowish white (cream), second segment roughened beneath with 
long loosely projecting dark brown scales, wholly overlaid with dark brown exteriorly except 
apical margin; terminal segment with a dark brown medial line ventrally. Head and thorax light 
yellow (pale buttercup yellow), lower part of frons, margin of eye and area around antenna dark 
brown ; tegula dark brown (negro) ; a sprinkling of brown scales at middle of thorax. Antenna 
yellowish brown, paler apically, suffused with greyish basally and overlaid with brownish scales 
at base; scape dark brown. Fore wing ground colour yellowish white, overlaid with yellowish 
brown except along inner margin (dorsum); cell and basal areas of wing suffused pale greyish; 
a blackish brown dash at f in phcal fold, a slightly heavier dot-like dash at end of cell ; a sprinkling 
of dark brown scales in plical fold, heaviest towards base; a more conspicuous irrorate longitudinal 
band of dark brown scales beginning between first stigma and costa, enclosing second stigma and 
reaching to apex and continuing into apical cilia; cilia otherwise yellowish white suffused with 
greyish, an admixture of brownish scales scattered along terminal margin and around apex. 
Hind wing greyish beige; cilia cream. 

Male genitalia (Figs. 122, 123). 

Congo: Lusinga, 1,760 m., 10.1v.1947 (G. F. de Witte), 1 holotype, 15 same locality data, 
various dates iii and iv.1947. Holotype and 10 paratypes in the Institut des Pares Nationaux, 
Brussels; 5 paratypes in the British Museum. 

This species is the only representative of the large composite genus Limnaecia known from the 
Ruwenzori region. It belongs in the L. eretmota species-group and may be distinguished by the 
more sombre coloration from other species of this group known to me. 



Diophila bathrota (Meyrick) comb.n. 
Mompha bathrota Meyrick, 191 1, Ann. Transv. Mus., 3 : 71. 
Diophila claricoma Meyrick, 1937, Exot. Microlepidopt., 5 : 149. Syn.n. 
ruwenzori: Ibanda, 4,700 ft. {Fletcher), 5 $ $. 

UGANDA Bundibugyo, 3,440 ft. (Fletcher), 7 £ $; Semliki Forest, 2,850 ft. (Fletcher), 5 S $• 

The above new synonymy has been established following an examination of type material in 
the British Museum. 

The type female of claricoma from Uganda is identical with the Ruwenzori specimens in 
coloration, and with them differs from the type female of bathrota from Transvaal, which has 
much lighter coloration. The abdomen of claricoma is missing but the genitalia of the type of 
bathrota (slide J.F.G.C. 4529) have been compared with the Ruwenzori specimens. 

The difference in coloration may indicate that the E. African material belongs to a distinct 
subspecies, for which the name claricoma could be used, but more material from the Transvaal is 
needed for study in order to establish this. 

Distribution: Transvaal and Uganda. 


Meridarchis mesosticha sp.n. (Figures 28, 124, 125) 

o 20 mm. Labial palpus white, second segment diffusedly irrorate blackish exteriorly except at 
apex. Head, thorax and tegula white, frons narrowly bordered with blackish around compound 
eye, posterior half of thorax suffused greyish, tegula similarly suffused, more strongly towards 
base. Antennal scape white, flagellum whitish-grey; ciliations about 3 times width of shaft. Fore 
wing white, partially suffused with dull ochraceous-buff mixed with fuscous, partially sprinkled 
with dark fuscous; a heavy blackish basal fascia inward-oblique from costa at 5, outer edge 
direct to inner margin (dorsum) and fringed with roughened white-tipped scales, fascia weak 
or obsolescent on inner margin at base; a thick blackish V-shaped marking with one side arising 
from costa at \, the other at f, both gradually thickening to converge a little beyond middle of 
wing, enclosing two blackish dots on costa; a third costal dot midway between marking and 
apex, below this dot weak blackish infuscation forming a diffuse narrow band direct to tornus; a 
circular whitish tuft in cell a little before middle adjacent to triangular marking, a second smaller 
tuft near costa, two similar less prominent tufts, one on each side of plical fold a little before 
raised scales forming two not very prominent contiguous tufts in outer margin of costal marking, 
one near middle, the other at apex of marking; cilia greyish, basal half slightly darker, apices 
often whitish producing a weak irrorate effect. Hind wing whitish grey; cilia pale buff, shaded 
greyish around apex and along termen except for narrow marginal line bordering wing. 
Male genitalia (Figs. 124, 125). 

ruwenzori: Nyinabitaba, 8,650 ft., 7-13.vii.1952 (Fletcher), 1 <J, holotypc, genitalia 
slide 8250. 

Near M. phaeodelta Meyrick, but readily distinguished by the strong black basal fascia on the 
fore wing. 


Ruwenzofi Expedition, 1952. Volume II, Number 12 

Carposina euschema sp.n. (Figures 29, 30, 126-130) 

c? § 20-31 mm. Labial palpus in male about i\ times as long as width of eye; in female about %\ 
times width of eye, second segment being greatly extended; white, second segment suffused 
fuscous-black exteriorly except at apex, terminal segment with fuscous-black medial annulus. 
Head whitish mixed with greyish or fuscous, most strongly on vertex and laterally behind 
antenna. Thorax white, almost wholly overlaid with fuscous-black except for diffuse anterior 
band; tegula fuscous-black, tipped with white. Antenna and scape whitish, suffused greyish 
fuscous, sometimes strongly, scape and other basal segments marked with fuscous-black; male 
ciliations about 3 times width of shaft; female with very short fine ciliations. Fore wing whitish, 
suffusedly mixed with fuscous which is variable and in some examples very dark; a weak sprink- 
ling of ochraceous-buff scales most evident in examples having light coloration; a fuscous or 
fuscous-black basal fascia inward-oblique from costa, sometimes black along distal margin, edged 
distally with white, weak or incomplete at base of dorsum (inner margin) ; infuscation heavier in 
cell area beyond middle and showing semblance of a medial fascia, edged outwardly with four 
black discal dots of raised scales, each dot ringed with white, the first three dots arranged in a 
straight line from middle of costa obliquely to end of cell, fourth dot coalescent and basad of 
third dot; two, sometimes three, similar dots at middle of wing along inner margin of infuscate 
area, these preceded by three similar dots extending obliquely inward across wing, two coalescent 
near costa, tbird situated near dorsum; six diffuse blackish spots on costa from \ to near apex; 
similar small neural spots along termen and around tornus; cilia whitish or grey, basal half barred 
with fuscous, apical \ tipped with fuscous. Hind wing whitish grey; cilia ochreous whitish, 
shaded with fuscous medially and apically. In the holotype and several paratypes the discal area 
of the fore wing is free of infuscation, leaving a large whitish patch. 

Male genitalia (Figs. 126-128). In the aedeagus of the type, Fig. 127, the large subapical spine- 
like seta is missing; it is shown in a paratype, Fig. 128. 

Female genitalia (Figs. 129, 130). Bursa copulatrix without signum. 

ruwenzori: Nyamaleju, 10,530 ft., 14-19. vii. 1952 (Fletcher), 4 including holotype, 
genitalia slide 8720, 5 $, including allotype, genitalia slide 8752; Bigo, 11,400 ft., 20-22.vii.1952 
(Fletcher), 1 <$, 1 $; Lake Bujuku, 13,050 ft., 22-28.vii.1952 (Fletcher), 1 

In size this species is similar to C. scieroto.xa Meyrick, of which I have not seen examples, but 
otherwise differs superficially in the distinctive dots ringed with white on the fore wing, and in 
the shorter antennal ciliations in the male, which according to the original description of 
scierotoxa are six times the width of the antennal shaft in that species. 

Carposina mesophaea sp.n. (Figures 31, 32, 1 31-13 5) 

o* 14 mm., $ 16-18 mm. Labial palpus in male about twice as long as width of eye; in female 
about 4X width of eye; second segment greatly lengthened; white, second segment suffused 
fuscous-black exteriorly except dorsal margin and apex; terminal segment with fuscous-black 
annulus medially. Head, chaetosema, thorax and tegula white ; mesothorax and base of tegula 
sometimes suffused fuscous-black. Scape white, antennal shaft white suffused fuscous. Fore wing 
white, a few scattered dark fuscous scales; a subtriangular blackish blotch occupying basal e of 
costa, a small roughened blackish spot between this and inner margin (dorsum) near base, well- 
defined in female, diffuse in male and connected to costal marking by fuscous suffusion; six 

Microlcpidoptera 103 

small blackish spots on costa from | to near apex; similar smaller blackish spots at ends of radial 
veins along termen and around tornus; 2 or 3 diffuse, sometimes confluent, elongate blackish 
spots transversely placed in disc at J; a heavy wedge-like blackish marking extending from end 
of cell to 4th black costal spot at f ; an ill-defined sinuous fascia from 5th costal spot to tornus 
(hardly discernible in holotype) ; cilia whitish-grey, basal half barred with fuscous along termen. 
Hind wing whitish-grey, darker distally; cilia whitish-grey, darker around apex. 

Male genitalia (Figs. 131, 132). 

Female genitalia (Figs. 133-135). 

ruwenzori: Nyinabitaba, 8,650 ft., 7-13.vh.1952 (Fletcher), 1 holotype, genitalia slide 
3270, 2 including allotype, genitalia shde 4223. 

The well-defined blackish discal marking, contrasting with the comparatively white general 
appearance of the fore wing, distinguishes this species from C. candace Meyrick, to which it is 
closely related. 

Carp o sin a poliophara sp.n. (Figures 33, 136, 137) 

18-20 mm. Labial palpus about if width of eye, white, second segment suffused fuscous-black 
exteriorly except apex; terminal segment with broad fuscous-black medial annulus. Head, thorax 
and tegula white; tegula greyish fuscous towards base, marked with fuscous-black at extreme 
base. Antennal flagellum whitish grey, ciliations about three times width of shaft; scape white. 
Fore wing white suffused with dull ochraceous-buff mixed with blackish; a weak inward-oblique 
fuscous mixed with ochraceous-buff basal fascia, incomplete proximally, strengthened with 
blackish admixture on costa; six small blackish spots spaced along costa from | to near apex, 
fifth and sixth spots slightly closer together; two raised ochraceous-buff suffused fuscous tufts 
below first costal spot arranged transversely across wing, first situated at middle in cell, second 
obhquely based on dorsal side of plical fold; a similar slightly larger tuft on costal side of fold a 
httle before middle of wing ; two equally large tufts close together and arranged transversely in 
discal area at end of cell, the one nearest costa preceded by a less prominent tuft on upper margin 
of cell; termen and tornus narrowly edged with fuscous-black, obscuring blackish neural spots; 
blackish irroration slightly heavier between fifth costal spot and tornus; a moderately heavy 
black dash below fourth costal spot between veins 9 and 10, and a similar black dash slightly 
basad on inner margin (dorsum) extending obliquely inward; cilia whitish suffused dark grey 
apically. Hind wing whitish grey; cilia white, suffused grey around apex. 

Male genitalia (Figs. 136, 137). 

ruwenzori: Nyinabitaba, 8,650 ft., 7-13.vii.1952 (Fletcher), 3 & including holotype, 
genitalia shde 8722. 

Near C. impavida Meyrick from which it may be distinguished by the shorter labial palpus 
and the absence of an extensive black discal patch. 

Eretmocera haemogastra Meyrick 
Eretmoccra haemogastra Meyrick, 1936, Exot. Microlepidopt., 5 : 62. 
Uganda: Semliki Forest, 2,850 ft. (Fletcher), 4 $, 4 $• 
Distribution: Congo. 

104 Ruwenzori Expedition, 1952. Volume II, Number 12 


Choreutis bjcrkandrella (Thunberg) 
Tinea bjcrkandrella Thunberg, 1784, Diss. Ent., 1 : 36, pi. 3, figs. 23, 24. 
ruwenzori: Jbanda, 4,700 ft. (Fletcher), 1 (J, 1 $. 
Distribution: Widespread and often common in both hemispheres. 

Choreutis pychnomochla sp.n. (Figures 34, 138-140) 

$ 1 5 mm. Labial palpus sordid white, brush of second segment mixed with sepia, an admixture 
of sepia on terminal segment strongest along upper margin. Head, thorax and tegula brownish 
grey (elephant skin colour), thorax darker posteriorly. Antenna greyish above, segments tipped 
with fuscous-black to about f, thence wholly fuscous-black to apex; wholly whitish at base 
ventrally, including scape, otherwise tipped with whitish to near apex. Fore wing brown, a 
strong whitish fascia from costa at J, inner margin well-defined, inward-oblique, slightly con- 
cave, outer margin diffuse, less oblique, outwardly dilated at middle, containing a sprinkling of 
brownish scales most evident towards inner margin (dorsum) ; basal area mixed with dark 
brown; an irrorate ferruginous streak from base below costa to fascia; a shorter less pronounced 
streak in plical fold; a diffuse narrow whitish fascia from costa at f, outward-curved at middle 
(discal area), excavation thus formed containing blackish scales; margin of costa between fasciae 
broadly suffused with dark brown; a sprinkling of blackish scales in space between fasciae 
extending from blackish patch in disc to inner margin; a heavy sprinkling of bluish grey scales 
in distal area, especially around apex and along termen to tornus; termen and apex outlined with 
ferruginous; cilia greyish brown, a dark brown basal line, a faint whitish medial line along 
termen. Hind wing fuscous; cilia brownish grey, a dark brown sub-basal line. 
Female genitalia (Figs. 138-140). 

ruwenzori: Nyamaleju, 10,530 ft., 14-19. vii.1952 (Fletcher), 1 holotype, genitalia slide 

A species of such distinctive appearance as to justify description from a solitary female. Similar 
in size to the Abyssinian species C. argyrastra Meyrick, of which it appears to be a near relative, 
but with much lighter general coloration in the fore wing and at once distinguished from this 
and other related species by the broad whitish fascia at 3. 

Choreutis agelasta sp.n. (Figures 35, 141-145) 

cT $ 13-15 mm. Labial palpus basal segment sordid white; second segment fuscous, irrorate with 
whitish dorso-medially and laterally towards apex; ventral brush fuscous-black. Head fuscous 
or greyish fuscous. Antenna fuscous, segments with raised scales at apices, ciliations fasciculate 
in both sexes, cilia approximately twice width of shaft in male, less than width of shaft in female. 
Thorax and tegula khaki with bronzy sheen. Fore wing khaki with bronzy sheen, without 
markings except for indication of a small whitish costal patch at about f; an admixture of 
ochreous scales around apex and along termen; cilia dark grey, a moderately broad basal band, 
overlaid in places by ochreous scales from termen; a thin whitish medial line, sometimes inter- 
rupted along termen. Hind wing grey, suffused brownish towards apex and in marginal areas; 

Microlepidoptcra I0 5 

cilia grey, a moderately broad dark sub-basal line edged on each side along termen by a thin 
whitish line. 

Male genitalia (Figs. 141, 142). 

Female genitalia (Figs. 143-145). 

ruwenzori: Bigo, 11,400 ft., 20-vii.i952 (Fletcher), 3 <J, including holotype, genitalia slide 
10494, 1 $, allotype, genitalia slide 10523; Nyamaleju, 10,530 ft., 14-19.vii.1952 (Fletcher), 6 <J. 

Near C. inspirata Meyrick, but distinguished from this and other members of the genus by 
the almost unicolorous appearance of the fore wing. 

Glyphipterix cfoalcotypa sp.n. (Figures 36, 146-149) 

ct $ 12-16 mm. Labial palpus light buff or whitish, suffused with fuscous below. Head, thorax 
and tegula light buff diffusedly irrorate with fuscous, fuscous irroration variable, in some examples 
dense and heavy especially on head. Antemia fuscous, in male finely ciliate, roughened in both 
sexes but more so in male, with raised scale-tippets at apices of segments. Fore wing light buff 
diffusedly irrorate with fuscous, more heavily in some examples than others; costal margin 
narrowly fuscous, containing five chalybeous dots, the first sometimes elongated, spaced along 
apical half of costa from middle to apex; thick silvery lines finely edged with dark fuscous 
arise from certain of these dots : from first dot near middle of costa extending transversely about 
^ across wing, from fourth dot extending, outward-curved, to termen near tornus, interrupted 
near middle of wing in some examples, from fifth dot (apical) extending along termen to middle, 
sometimes fusing with interrupted apical portion of line from fourth dot or with distal part of 
same line; in latter examples whole of termen appears edged with silver; short silvery blobs 
edged proximally with dark fuscous arise from second and third dots; a silvery line similar to 
those emitted by costal dots arises from middle of dorsum, curving gently distad to terminate 
beyond middle of wing near third costal dot; directly distad of this line is a narrow light yellow 
(butter yellow) fascia reaching across wing to costal margin, enclosing first and second chalybeous 
dots and extending apicad as far as third dot; space between silvery lines emitted by fourth and 
fifth costal dots pale yellow; fourth silver line bordered with pale yellow along inner margin; 
a short thick silvery line below costa at about I in proximal margin of fascia; a black flash sparsely 
sprinkled with yellow extending inward-oblique from tornus to silvery line from dorsum, with 
three or four embossed cupreous bands; cilia light buff, irrorate with fuscous, a broad light 
yellow basal band along termen from apex to tornus. Hind wing light grey; cilia similar, a cream 
buff basal shade along termen and inner margin, a diffuse grey sub-basal line along termen. 

Male genitalia (Figs. 146, 147). 

Female genitalia (Figs. 148, 149). 

Congo: Lusinga, 1,760 m., 10.iv.1947 (G. F. de Witte), 1 holotype; R. Mubalc, 1,480 m., 
1-20.V.1947 (G. F. de Witte), 9 $, 10$, including allotype; Buye-Bala at g. Muye (af. dr. Lufira), 
1,750 m., 25-3 1. iii. 1948 (G. F. de Witte), 1 6"; Kankunda am. g. Lupiala, 1,300 m., 14-20.xi.1947 
(G. F. de Wittte), 1 <J. 

Holotype, allotype and 15 paratypes in the Institut des Pares Nationaux, Brussels; 5 paratypes 
in the British Museum. 

This species is near G. ditiorana (Walker) which occurs in South Africa. The two species may 
be separated by differences in the silvery lines arising from the apical and preapical costal dots 

106 Ruwenzori Expedition, 1952. Volume II, Number 12 

on the fore wing, in ditiorana the line from the apical dot extends along the termen to tornus, 
in chalcotypa it normally reaches only to middle of termen; the preapical line in ditiorana extends 
to middle of wing, in chalcotypa this line normally traverses the wing to termen above tornus. 

Glyphipterix plagiographa sp.n. (Figures 37, 150, 151) 

o* 11 mm. Labial palpus pastel yellow, second segment with sepia medial and basal annuli; 
terminal segment dark sepia below and at apex, a somewhat diffuse sepia annulus at base. Head, 
thorax and tegula dull silvery cinereous, head narrowly edged with pastel yellow anteriorly 
from base of antenna ; fringing scales (eyelashes) along posterior margin of eye pastel yellow ; a 
trace of pale yellow in tegula. Antenna greyish sepia. Fore wing dark greyish sepia mixed with 
earth colour, a short transverse yellow bar on costa at I, a similar much smaller deeper yellow 
strigula a little apicad, followed by six transverse whitish streaks, partially edged with blackish, 
spaced along costa to near apex, the basal four slightly outward-oblique, each streak white and 
comparatively conspicuous at costa, all except second, which is shorter than the others, strongly 
violaceous inwardly ; a small blackish spot below costa close to apex ; a small elongate blackish 
patch in disc surrounded by several short, broken, transverse chains of violaceous scales reaching to 
inner margin; termen diffusely edged with blackish sepia, indented below apex; cilia whitish, 
a blackish sepia basal line. Hind wing fuscous; cilia matching, paler apically. 
Male genitalia (Figs. 150, 151). 

ruwenzori: Nyinabitaba, 8,650 ft., 7-13.vii.1952 (Fletcher), 1 holotype, genitalia slide 

Similar to G. gemmatclla (Walker) but lacking the broad yellow sub-basal fascia on the fore 

Glyphipterix gemmatella (Walker) 
Gelechia gemmatella Walker, 1864, Cat. Lcp. Ins. B.M., 30 : 1022. 
Uganda: Semliki Forest, 2,850 ft. (Fletcher), 2 <$. 
Distribution: Sierra Leone and French Congo. 


Elachista oritropha sp.n. (Figures 38, 39, 152-156) 

ct $ 10-11 mm. Labial palpus smooth-scaled, dark silver varying to smoke brown, whitish 
interiorly. Front of head (frons) smooth-scaled, somewhat shining, whitish or whitish-grey; 
crown with loosely appressed whitish or greyish fuscous scales. Antenna dark silver varying to 
smoke brown. Thorax and tegula silver varying to smoke brown. Fore wing light silver, suffused 
smoke brown or light sepia, suffusion usually weak distally so that the silvery ground colour is 
apparent, becoming stronger towards base of wing, and in some examples very heavy; a some- 
what diffuse broad sepia streak from below costa at about 3 extending very obliquely to termen 
below costa, slightly thickened with a small darker coloured discal dash between cell and apex; 
a similar parallel streak in plical fold from near base to a little beyond middle of wing; cilia greyish 
brown. Hind wing greyish silver ; cilia greyish brown. 

Micwlepidoptera IO y 

This is a variable species and the above description is of the commonest form among the 
material available and is considered typical. A form (Fig. 39) occurs in which the interior of the 
labial palpus, including apex of second segment, the whole of head and thorax medially, are 
pastel yellow ; and the fore wing is more heavily suffused with dark brown or fuscous, excepting 
along dorsal margin which is whitish, sometimes with an admixture of yellow, and in most 
examples with a short outward-oblique linear dash projecting from before middle; the dorsal 
streak terminating in a small similarly coloured blotch at tornus; a similar slightly smaller blotch 
obliquely above on costa at about f . 

Male genitalia (Figs. 152, 153). 

Female genitalia (Figs. 154-156). 

ruwenzori: Nyamaleju, 10,530 ft., 14-19.vh.1952 {Fletcher), 33 <J, including holotype, 
genitalia slide 10129, 7 including allotype, genitalia slide 9612; Bigo, 11,400 ft., 20-22. vii. 1952 
(Fletcher), 4 2 £; Lake Bujuku, 13,050 ft., 22-28. vii 1952 (Fletcher), 1 6*; Lamia Valley, 11,900 ft., 
30-3 1. vii. 1952 (Fletcher), 1^,1$; Lake Mahoma, 9,600 ft., 12.v11.1952 (Fletcher), 9 2 

A distinctive species by virtue of the wing pattern which differs from that of other known 
species of Elachista, and the only representative of this genus recorded from the alpine region 
of Ruwenzori. 


Scythris philorites sp.n. (Figures 40, 157-159) 

o* 10-13 mm. Labial palpus pale yellow, apical half of second segment and whole of terminal 
segment except base overlaid with fuscous exteriorly. Head, thorax and tegula bronze, head 
shiny, tip of tegula and medial area of thorax tinged purplish. Antenna fuscous-black. Fore wing 
fuscous-black, moderately sprinkled with whitish scales along costal margin and in apical area; 
a broad white streak from base along plical fold to tornus, slightly constricted at middle and 
dilate near tornus ; cilia dark grey, those below apex to near middle of termen pale yellow apically. 
Hind wing dark grey, cilia concolorous. At the base of the abdomen a pair of hair-pencils arise 
dorso-laterally on the first somite, as shown in Fig. 159. "When not expanded these are concealed 
in longitudinal pockets or folds in the first and second pleurites. 
Male genitalia (Figs. 157, 158). 

ruwenzori: Nyamaleju, 10,530 ft., 14-19. vii. 1952 (Fletcher), 2 including holotype, 
genitalia slide 10137. 

A distinctive species, tentatively assigned to the composite genus Scythris and placed near S. 
aphanatma Meyrick which it resembles in size and general coloration. The two species may be 
separated by the absence of the white plical streak on the fore wing in aphanatma, and by the 
coloration of the head which is light yellow in aphanatwa, compared with bronze in philorites. 


Argyresthia lamiella sp.n. (Figures 41, 42, 160-166) 
6* $ 11-12 mm. Labial palpus greyish, apical halves of second and terminal segments suffused 
fuscous. Head whitish grey, roughened scales on crown and vertex light greyish ochrcous or dull 
ochraceous. Antenna whitish grey, flagellum with somewhat diffuse greyish fuscous annuli. 

108 Ritwenzori Expedition, 1952. Volume II, Number 12 

Thorax and tegula somewhat shining pale golden or coppery golden; thorax broadly suffused 
with whitish grey or grey medio-dorsally. Fore wing coloration and markings variable, male 
usually with markings more obscure than in female; ground coloration in male white suffused 
with a mixture of pale golden and fuscous, a series of compacted striae along costa from about ^; 
several diffuse ill-defined whitish patches along inner margin (dorsum), a sprinkling of whitish 
scales in apical area; female with ground colour whitish, a semblance of three golden-brown 
fasciae, first strong on inner margin at about \ forming a quadrate blotch, obsolete at middle of 
wing, reappearing on costa; second at about middle of inner margin, similar and parallel with 
first; third terminal, very narrow, not always discernible; costa basad of first fascia usually pale 
grey, a sprinkling of whitish scales around apex and along termen; cilia pale grey v/ith a weak 
golden tinge. Hind wing pale grey; cilia matching, with a weak golden tinge. 

Male genitalia (Figs. 160-163). 

Female genitalia (Figs. 164-166). 

ruwenzori: Lamia Valley, 11,900 ft., 30-3 i.vii. 1952 (Fletcher), 1 holotype, genitalia slide 
10456, 5 including allotype, genitalia slide 10452; Bigo, 11,400 ft., 20-22. vii. 1952 (Fletcher), 
2$, 1$; Nyamaleju, 10,530 ft., 14-19. vii. 1952 (Fletcher), 5 o\ 2$; Lake Mahoma, 9,600 ft., 12. vii. 
1952 (Fletcher), 1 o\ 1 

In general appearance this species is reminiscent of the European A. goedartella (Linnaeus), but 
the two are not likely to be confused since they are allopatric. The only other Argyresthiid 
belonging to this genus known from E. Africa is A. liparodes Meyrick, which is readily recogniz- 
able from lamiella by the whitish head and labial palpus, and the whiter general appearance of the 
fore wing which lacks the golden-brown fasciae of lamiella. 


Ethmia phricotypa sp.n. (Figures 43, 167, 168) 

o* 24 mm. Labial palpus black, second segment lightly sprinkled with whitish scales exteriorly 
and below, narrowly ringed with white apically; terminal segment with white submedian and 
apical annuli. Head with face purplish black, crown white. Antenna purplish black, scape white 
below. Thorax and tegula white, a purplish black quadrate blotch at base of tegula, a similar 
marking medially on prothorax, a third smaller marking on metathorax. Fore wing purplish 
black except on inner margin along which extends a narrow white streak the inner edge of which 
makes strong incursions into the black area as follows : near base, at J, this closely followed by a 
slight indentation, and before tornus; a large white post-tornal blotch extends into cilia up to 
middle of termen; cilia beyond this point purplish black, somewhat brownish around apex; 
costal margin of wing sparsely sprinkled with sordid white. Hind wing butter yellow basally, 
becoming greyish distally; a pecten of long greyish yellow hair-like scales arises on basal half of 
subcostal vein. Fore and middle legs black; tibia of middle leg ringed with white at middle, and 
bearing small white external spots at base and apex; tibia of fore leg similarly but less strongly 
marked, whitish annuli at base and apex of first (basal) tarsal segment; hind leg deep butter yellow 
becoming blackish towards apex of first tarsal segment, tarsal segments marked with white apical 
bars dorsally. 

Male genitalia (Figs. 167, 168). 

Uganda: Bugoye, 4,500 ft., 5-10.ix.1952 (Fletcher), 1 o\ holotype, genitalia slide 10474. 

Microlepidoptera log 

Near E. iphicrates Meyrick from which it is readily separated by the solid black coloration of 
the costal half of the fore wing and by the yellowish coloration of the hind wing. 


Coleophora tacera sp.n. (Figures 44, 169-171) 

S § 1 1-1 5 mm. Labial palpus with terminal segment slightly larger than second ; projecting apical 
tuft on second weakly developed, very short; sepia, in male usually wholly overlaid with light 
yellow interiorly, but apex remaining sepia in some examples; terminal segment sometimes 
suffused with light yellow or partially suffused to near apex; in female second segment suffused 
yellowish white at base. Head shining light golden-yellow. Antenna greyish sepia, apical \ 
whitish; scape comparatively short and simple, without pecten; scape and basal segments of 
flagellum sometimes with a weak golden sheen. Thorax and tegula greyish golden-yellow. Fore 
wing dark grey with deep golden sheen, becoming more yellow towards apex, reddish yellow or 
buttercup yellow at apex; some examples weakly violaceous distally; cilia yellowish (matching 
wing) around apex, grey along termen. Hind wing dark grey, cilia paler. 

Male genitalia (Fig. 169). 

Female genitalia (Figs. 170, 171). 

euwenzori: Bigo, 11,400 ft., 20-22. vii. 1952 [Fletcher), 3 $, including holotype, genitalia 
slide 10139, 1 $, allotype, genitalia slide 9617; Nyamaleju, 10,530 ft., 14.vi1.1952 [Fletcher), 5 c?> 
4$; Lake Mahoma, 9,600 ft., 12.vii.1952 [Fletcher), 1 

Related to the palaearctic C. idaeclla Hofmann and its Vaccinium-keAmg allies, but distinguished 
from these and other members of the genus by the yellowish coloration in the apical area of the 
fore wing, which appears to be peculiar to tacera. 


Conopomorpha fustigera (Meyrick) 
Acrocercops fustigera Meyrick, 1928, Exot. Microlepidopt., 3 : 408. 
Uganda: Bundibugyo, 3,440 ft. [Fletcher), 1 5*. 
Distribution: Uganda (Kampala) and South Africa (Durban). 

Acrocercops orianassa Meyrick 
Acrocercops orianassa Meyrick, 1932, Trans, ent. Soc. Lond., 80 : 116. 
ruwenzori: Nyinabitaba, 8,650 ft. [Fletcher), 1 
Distribution: Abyssinia. 

Acrocercops chenopa Meyrick 
Acrocercops chenopa Meyrick, 1932, Exot. Microlepidopt., 4 : 270. 
ruwenzori: Mahoma River, 6,700 ft. [Fletcher), 1 <3\ 1 $• 
Distribution: Uganda (Ruwenzori). 


Ruwenzori Expedition, 1952. Volume II, Number 12 

Metriochroa celidota sp.n. (Figures 45, 172-175) 

o* $ 7-9 mm. Labial palpus sordid white, suffused with greyish or blackish fuscous, usually more 
strongly exteriorly. Head greyish with a strong gloss, crown darker posteriorly. Thorax and 
tegula greyish or fuscous mixed with greyish. Antenna and scape dark greyish fuscous; apical 
fifth of antenna whitish. Fore wing blackish fuscous, somewhat lightened by an admixture of 
greyish basad of postmedial fascia; two silvery- white transverse fasciae, first at about J poorly 
defined and partly obliterated, especially towards inner margin (dorsum) by a suffusion of greyish 
fuscous; second fascia at about f, comparatively bold and well-defined, free from extraneous 
suffusion, extending a little into cilia at tornus, inner edge almost straight and slightly inward- 
oblique from costa, outer edge shallowly concave; an obscure partial fascia or transverse elongate 
diffuse white patch near costa a little before middle, absent in female; cilia dark grey, paler around 
termen and inner margin. Hind wing and cilia grey. Legs wholly grey, depth of colour varying 
between specimens. 

Male genitalia (Figs. 172, 173). 

Female genitalia (Figs. 174, 175). 

ruwenzori: Nyamaleju, 10,530 ft., 14-19. vii.1952 (Fletcher), 4 including holotype, 
genitalia slide 9623; 1 §, allotype, slide 10453. 

Near M. argyrocelis Vari, described from S. Rhodesia, but at once distinguished superficially 
by difference in arrangement of the white markings of the fore wing, argyrocelis having small 
broken markings on the costa and tornus in place of the complete transverse postmedial fascia 
in celidota. 

Caloptilia pachyspila sp.n. (Figures 46-48, 176-180) 

o* 9-12 mm. Labial palpus dark grey. Head varying from wholly whitish to dark grey, frons often 
paler than crown. Thorax and tegula dark grey. Antenna dark grey, a broad whitish subapical band. 
Fore wing pattern variable, type (Fig. 46) and one paratype as follows : ground colour fuscous- 
black diffusedly mixed with greyish, marked with three white fasciae, the first sub-basal, very 
diffuse and comparatively inconspicuous ; second a little before middle, comparatively bold and 
broad, directly transverse and with well-defined margins; third at about f, outer edge directly 
transverse, inner edge slightly incurved from costa; a weak bronzy sheen covers the whole wing 
and is also faintly discernible on the thorax and crown of head; of the remaining paratypes, five 
have generally lighter ground coloration than the type, the coloration of the head is also lighter 
(whitish), but the fore wing lacks the bold white fasciae, these being represented by small white 
markings, varying in size and number, along the costal and dorsal margins and in the apical 
area, as shown in Fig. 47; in these paratypes the bronzy sheen is evident; in two paratypes from 
Mt. Stanley (Fig. 48) and Balirungi R. the fore wing is a uniform fuscous-black diffusedly mixed 
with greyish, the white markings being entirely absent, and in only the Balirungi R. specimen is 
there a trace of the bronzy sheen. Hind wing and cilia grey. 

Male genitalia (Figs. 176-180). In the holotype, Fig. 176, the prong projecting from the ventral 
margin of the valva a little basad of the apical flange is vestigial but in some examples examined 
was found to be more strongly developed as shown in Fig. 178. In Fig. 180 are shown the expans- 
ible hair-tufts arising laterally on the Vllth somite. 

Microlepidoptera hi 

ruwenzori: Lake Mahoma, 9,600 ft., 12.vii.1952 {Fletcher), 2 q*, including holotype, genitalia 
slide 8937; Balirungi River, 11,200 ft., i.vii.1952 (Fletcher), 1 Bigo, 11,400 ft., 20-22. vii.1952 
(Fletcher), 3 6*; Mt. Stanley, 14,500 ft., 26.vii.1952 (Fletcher), 1 

The remarkably wide variation in the superficial appearance of pachyspila does not appear 
from the material examined to be related to locality or elevation. The sombre general coloration 
and, when present, the white markings of the fore wing, distinguish this species from the 
species which follows, C.janeae Bradley, next which it is tentatively placed. 

Caloptilia janeae sp.n. (Figures 49, 50, 1 81-186) 

cT 9 16-21 mm. Labial palpus blackish sepia, apex of terminal segment, base and underside of 
second segment overlaid with pale sulphur yellow. Head blackish sepia, mixed with pale sulphur 
yellow laterally. Antenna dark grey. Thorax and tegula blackish sepia, thorax with a longitudinal 
sulphur yellow lateral streak, tegula tipped with pale sulphur yellow. Fore wing sulphur yellow, 
markings variable, consisting mostly of small, irregular, scattered blackish sepia dots and patches, 
sometimes confluent medially; cilia brownish grey from apex to inner margin, an admixture of 
dark sepia along termen, sometimes a sprinkling of sulphur yellow at base along termen; indian 
yellow above apex. Hind wing grey, cilia brownish grey. 

Two of the paratypes, a male and female from the type locality, represent a form in which the 
sulphur yellow coloration of the fore wing is replaced with light brown, the male having the 
dark markings greatly reduced and less intense than in the typical form, the female lacking these 
markings altogether and having the fore wing an almost uniform dark brown. 

Male genitalia (Figs. 1 81-183). 

Female gemtalia (Figs. 184-186). 

ruwenzori: Nyamaleju, 10,530 ft., 14-19.vii.1952 (Fletcher), 4 <$, including holotype, 
genitalia slide 10131, 2 $, including allotype, genitalia slide 9586; Bigo, 11,400 ft., 20-22. vii.1952 
(Fletcher), 3 Lake Mahoma, 9,600 ft., 12.vii.1952 (Fletcher), 1 <J. 

The sulphur yellow coloration and the intricate pattern of broken blackish sepia markings of 
the fore wing in the typical form distinguish this species from C. pachyspila and other members 
of the genus. 


Phyllocnistis loxosticha sp.n. (Figures 51, 187-190) 
6* 9 7 mm. Labial palpus, head, thorax and tegula white, a trace of yellowish sheen on palpus and 
head; thorax broadly suffused with dark olive brown medially; tegula narrowly edged with 
dark olive brown along outer margin. Antenna white; a longitudinal olive brown dorsal line 
from base of scape to apex of flagellum. Fore wing white, a weak yellowish sheen sometimes 
evident; two thick parallel median longitudinal dark olive brown lines from base, upper (costad) 
reaching to middle, lower to §, enclosed space greyish pale yellow; a dark olive brown line from 
costa at I, outward-oblique from costa to middle thence inward-oblique to inner margin (dor- 
sum), followed by two similar streaks from costa, originating in cilia, and preceded by a similar 
but stronger and well-defined very oblique line from middle of costa, terminating at end of lower 
ohve brown medial line, all these lines from costa inwardly edged with a broad suffusion of pale 

112 Ruwenzori Expedition, 1952. Volume II, Number 12 

yellow; an elongate jet black apical clash; cilia white, basal half suffused pale yellow along termen, 
a straight dark olive brown medial line from costa to a point level with inner margin (dorsum) ; 
four blackish diverging bars in apical cilia arising from apex of wing. Hind wing greyish white; 
cilia white. Legs sordid white; tibia and tarsus of fore leg suffused fuscous-black interiorly; 
hind tibia with comb of dorsal hairs. 

Male genitalia (Figs. 187, 188). 
Female genitalia (Figs. 189, 190). 

ruwenzori: Mahoma River, 6,700 ft., 13-16.viii.1952 (Fletcher), 3 <$, including holotype, 
genitalia slide 10145, 2 9, including allotype, genitalia slide 10526. 

Allied to the palaearctic P. saligtia Zeller, but differing superficially in the very much more 
oblique linear marking from the middle of costa, and in the elongate black apical dash which in 
saligna is in the form of a round dot. 


Epermenia oriplanta sp.n. (Figures 52, 53, 191-195) 

o* $ 13-27 mm. Both sexes variable in size and appearance; in the type series the majority measure 
about 22-24 mm., the largest in the series being the holotype male, and the smallest a paratype 
male with the same data. Labial palpus warm buff or ochraceous-buff, infuscate exteriorly; 
terminal segment and apical part of second segment sometimes overlaid with fuscous-black. 
Head warm buff or ochraceous-buff, variably suffused greyish, fuscous or fuscous-black; frons 
sometimes with whitish admixture. Thorax and tegula warm buff or ochraceous-buff mixed or 
wholly overlaid with greyish, fuscous or fuscous-black; frons sometimes with whitish admixture. 
Thorax and tegula warm buff or ochraceous-buff mixed or wholly overlaid with greyish, fuscous 
or fuscous-black; tegula frequently tipped with white scales. Antenna in female minutely ciliate 
along anterior margin to near base, basal segments, dorsal and posterior surfaces of flagellum clad 
with greyish fuscous or fuscous scales, usually darker towards apex of antemia, usually a trace of 
ochraceous admixture in basal segments, individual segments often lighter proximally, scape 
fuscous-black dorsally with a trace of ochraceous admixture, underside of scape and basal segments 
of flagellum whitish cream; antennal flagellum in male wholly ciliate, ciliations about \ width 
of shaft, shaft thickened beyond base, gradually tapering apically, scape and basal segments clad 
with fuscous scales with variable ochraceous admixture, remainder of segments except near apex 
each spotted posteriorly with a small scallop-like tuft of whitish scales. Fore wing ground colour 
white, overlaid with variable fuscous-black irroration, a variable admixture of ochraceous-tawny, 
in some examples this is very strong and forms diffuse patches especially in middle and discal 
areas of wing ; inner margin (dorsum) bordered with white to beyond middle, a raised blackish 
scale-tooth at about J, a smaller tuft beyond middle at end of white border; sometimes a few 
white scales partly encircling a black discal dot at end of cell, less frequently a sprinkling of white 
scales extending into cell area; cilia light buff, a blackish medial line along termen, cilia apicad 
of tornus to near middle of termen suffused fuscous. In allotype female and four paratype males 
the ochraceous-tawny coloration is obliterated by heavy fuscous and fuscous-black suffusion; in 

Microlepidoptera 113 

the allotype the blackish coloration is exceptionally heavy and forms a narrow inward-oblique 
blackish mixed with ochreous medial fascia. Hind wing silvery grey; cilia light buff. 

Male genitalia (Figs. 191, 192). 

Female genitalia (Figs. 193-195). 

ruwenzori: Lake Bukuju, 13,050 ft., 22-28.vh.1952 (Fletcher), 6 6*, including holotype, 
genitalia slide 8266, 1 allotype, slide 4221; Bigo, 11,400 ft., 20-22.vii.1952 (Fletcher), 1^,1$; 
Lamia Valley, 11,900 ft., 30-3i.vii.i952 (Fletcher), 1 $\ Nyamaleju 10,530 ft., 19.vii.1952 
(Fletcher), 5 Lake Mahoma, 9,600 ft., 12.vii.1952 (Fletcher), 1 q\ 

This species is apparently a close relative of E. epirrhicna Meyrick, described from the Pare 
National Albert from one specimen, which I have not seen. E. oriptanta differs superficially from 
epirrhicna which, according to the original description, does not have ochraceous-tawny coloration 
in the fore wing and has a basal patch of dark fuscous suffusion, and has the thorax white mixed 
with grey. 


Plutella symmorpha sp.n. (Figures 54, 196-198) 

o 16 mm. Labial palpus pale cream-buff, suffused fuscous-black exteriorly; apical tuft of second 
segment short. Head and antennal scape pale cream-buff mixed with cartridge buff, weakly 
suffused with greyish. Antennal flagellum cartridge buff, annulate with ochraceous-buff Thorax 
and tegula pale cream-buff heavily overlaid with fuscous-black. Fore wing greyish fuscous, 
variably irrorate or suffused with fuscous-black; several more or less distinctly developed longi- 
tudinal blackish lines radiating from cell area in costal half of wing; dorsal area as far as fold 
relatively free of dark infuscation, a diffuse bisinuate ochraceous-buff streak extending along 
plical fold to inner margin (dorsum), more or less edged above with blackish; a slender wedge- 
shaped ochraceous-buff streak from middle of termen to middle of wing ; blackish pre-apical 
strigulae along costa interspersed with ochraceous-buff; termen diffusedly edged with blackish; 
cilia cartridge buff mixed with ochraceous-buff, a diffuse sub-basal line, a few cilia tipped with 
blackish. Hind wing light grey; cilia whitish grey, with a grey sub-basal shade. 
Male genitalia (Figs. 196-198). 

kenya: Aberdare Range, Mt. Kinango, 8,000 ft. (Edwards), 1 6\ holotype, genitalia slide 8268. 

Related and superficially similar to the Abyssinian species P. stichocentra Meyrick, but disting- 
uished by its smaller size, the general coloration of the fore wing, which in stichocentra is distinctly 
more grey along the costa, and the distribution of the blackish irroration, which in stichocentra is 
less concentrated and does not form the almost unbroken radiating lines as in symmorpha. 

Plutella orosoma Meyrick 
Plutella orosoma Meyrick, 1932, Exot. Microlepidopt., 4 : 228. 

ruwenzori: Lake Bujuku, 13,050 ft. (Fletcher), 2 3; Lamia Valley, 11,900 ft. (Fletcher), 1 $; 
Bigo, 11,400 ft. (Fletcher), 15 <J, 4 9; Nyamaleju, 10,530 ft. (Fletcher), 4 c?, 2$; Lake Mahoma, 
9,600 ft. (Fletcher), 1 o*. 

Distribution: Uganda (Ruwenzori). 


Ruwenzori Expedition, 1952. Volume II, Number 12 

A truly montane species of Plutellid having greatly developed wing expanse and occurring 
mainly at higher elevations in the ericaceous zone and extending into the alpine belt. The fresh 
material shows evidence of considerable superficial variation within the species. In the typical 
form the dorsal markings of the fore wing are well-defined and the inner and terminal margins 
are mixed with whitish; in some examples the margins are diffuse to varying degrees and may 
sometimes be mixed with ochreous. 

P. orosoma appears to be a near relative of the Abyssinian species P. oxylopha Meyrick and the 
European species P. iucariwtella Steudal, but most remarkable is that it seems to be more nearly 
related to P. culminate Meyrick, which occurs in South America. A comparison of the genitalia 
of the type female of culminata, which is in the British Museum, with those of orosoma shows 
them to be structurally very similar. 

Acrolepia nephelota sp.n. (Figures 55, 199-203) 

o* $ 11-12 mm. Labial palpus yellowish- white, second segment suffused fuscous below; terminal 
segment with broad fuscous sub-basal and apical annuli. Head, thorax and tegula fuscous, with 
a weak dark purplish sheen ; front of head with variable admixture of yellowish- white ; tegula 
becoming yellowish-white towards tip; patagia yellowish-white or ochreous. Antenna greyish 
fuscous; scape fuscous. Fore wing greyish fuscous irregularly and diffusely strigulate with blackish; 
several poorly defined thick fuscous-black strigulae along costa; a moderately heavy sprinkling 
of white scales forming a whitish patch in distal area below costa at about §; a thin sprinkling of 
white scales at apex and along termen, a few on costa immediately before apex sometimes forming 
a small whitish dot at about f, also a few on inner margin (dorsum) before tornus and at middle; 
cilia greyish fuscous. Hind wing dark grey; cilia matching. 

Male genitalia (Figs. 199, 200). 

Female genitalia (Figs. 201-203). 

ruwenzori: Lake Mahoma, 9, 600 ft., 12. v. 1952 (Fletcher), 1 holotype, genitalia slide 10458; 
Nyamaleju, 10,530 ft., 14-19.vii.1952 (Fletcher), 2 including allotype, slide 9627. 

Near A. trapezopa Meyrick, described from Mt. Mlanje; distinguished by the darker coloration 
of the fore wing and absence of the subquadrate blotch present on the inner margin in trapezopa. 


Opogona chrysophthalma Meyrick 

Opogona chrysophthalma Meyrick, 1934, Exot. Microlepidopt., 4 : 462. 

Uganda: Entebbe, Lake Victoria, Bugonga Point, Cave No. 2 (Fletcher), 13 $ $. 
kenya : Nairobi (Edwards), 1 $. 

Distribution : Uganda and Kenya. 

Opogona dimidiatella Zeller 

Opogona dimidiatella Zeller, 1853, Byull. mosk. Obshch. Ispyt. Prir., 26 (2) : 507, pi. 4, figs. 13-16. 
Glyphypteryx dimidiatella Walker, 1864, Cat. Lep. Ins. B.M., 30 : 839. Syn.n. 

ruwenzori: Ibanda, 4,700 ft. (Fletcher), 1 
Uganda: Semliki Forest, 2,850 ft. (Fletcher), 1 1 

Microlepidoptera 115 

The above new synonymy has been established following an examination of the type of 
O. dimidiatella Zeller (a male, genitalia slide 1862) and the type of G. dimidiatella Walker (a female, 
slide 8975), which are both in the British Museum. 

Distribution : Indo-Malayan region, South and East Africa. 

Opogona scaeozona Meyrick 

Opogona scaeozona Meyrick, 1920, Voyage de Ch. Alluaud et R. Jeannel en Afrique Oriental, 2, 
Microlepidoptera, p. 94. 

ruwenzori: Mahoma River, 6,700 ft. (Fletcher), 3 $\ Nyinabitaba, Lower Bamboo zone, 
8,700-9,000 ft. {Fletcher), 4 

Distribution: Kenya. 


Tinissa poliophasma sp.n. (Figures 56, 204, 205) 

§ 19 mm. Labial palpus white, second segment narrowly suffused with dark sepia dorsally to near 
apex, an admixture of sepia in middle of brush exteriorly and interiorly; terminal segment with 
narrow sepia annulus medially. Head, thorax and tegula white, base of tegula suffused with beige. 
Antennal flagellum beige; scape white. Fore wing beige diffusely irrorate with slightly darker 
flecks giving the wing a somewhat mottled appearance; a white fascia at base, outer margin 
straight, inward-oblique from costa at about i, narrowly edged with dark beige; base of costa 
and extreme base of wing dark beige; a narrow irregular dark beige line from costa at middle of 
white basal fascia, inward-oblique towards base of inner margin (dorsum), strong and well- 
defined at costa becoming weak and obsolete near middle and beyond ; a pair of small indistinct 
strigulae on costa a little before middle; a white transverse spot on costa at f, a similar smaller 
spot at f ; small diffuse dark beige interneural spots around apex and along termen; an indistinct 
whitish spot on dorsum near middle, and a smaller pretornal spot; cilia beige, suffused blackish 
along termen. Hind wing greyish with a weak golden sheen, apex and termen thinly outlined 
with dark grey, an elliptical patch of white scales on basal half of costa (concealed by fore wing) ; 
cilia grey, suffused with light buff basally along margin of wing, apices white-tipped. 
Female genitalia (Figs. 204, 205). 

Uganda: Semliki Forest, 2,850 ft., 22.vi1i.-3.ix.1952 (Fletcher), 1 holotype, genitalia slide 

The conspicuous white basal fascia readily distinguishes this species from T. spauiastra Meyrick. 
described from Abyssinia, to which it appears to be most closely related. 

Polymnestra capnochalca Meyrick 
Polymnestra capnochalca Meyrick, 1932, Exot. Microlepidopt., 4 : 207. 
ruwenzori: Bigo, 11,400 ft. (Fletcher), 1 6*- 

Previously known only from the original two type specimens taken on Ruwenzori at 
10,000-12,000 ft. 
Distribution: Uganda (Ruwenzori). 


Ruwevzori Expedition, 1952. Volume II, Number 12 

Atabyria bucephala Snellen 

Atabyria bucephala Snellen, 1884, Tijdschr. Ent., 27 : 166, pi. 9, figs. 1, ia-b. 
Osphretica chomatias Meyrick, 1910, Trans, ent. Soc. Lond., 1910 : 475. 
Dcprcssaria rotundata Matsumura, 193 1, 6000 111. Ins. Japan, p. 1091, fig. 2248. 

Uganda: Semliki Forest, 2,850 ft. {Fletcher), 1 
ruwenzori: Mahoma River, 6,700 ft. (Fletcher), 1 

Distribution: S. Africa, India, China, Borneo, E. Siberia and, if the synonymy of rotundata is 
correct, Japan. 

Trapezoritis anisastra Meyrick 
Trapezoritis anisastra Meyrick, 1932, Exot. Microlepidopt., 4 : 212. 
ruwenzori: Nyamalejn, 10,530 ft. (Fletcher), 1 q\ 
Distribution: Uganda (Ruwenzori). 

Homalopsycha hyacinthopa Meyrick 
Homalopsycha hyacinthopa Meyrick, 1932, Exot. Microlepidopt., 4 : 208. 

ruwenzori: Namwamba Valley, 6,500 ft. (Edwards), 1 Namwamba Valley, 11-12,000 ft. 
(Edwards), 1 Lamia Valley, 11,900 ft. (Fletcher), 1 3*; Nyamgasani Valley, 12,500 ft. (Buxton), 
1 <J» 1 $; Portal River, 12,250-12,550 ft. (Fletcher), 2 Bujuku River, 12,550 ft. (Fletcher), 1 1 
Lake Bujuku, 13,050 ft. (Fletcher), 6 6*, 2 $; Stuhfman Pass, 13,500 ft. (Fletcher), 5 0*; Mt. Stanley, 
14,500 ft. (Fletcher), 1 

Uganda: Kilembe, 4,500 ft. (Edwards), 1 

This species has previously been known only from the type, a male collected at Bujuku at 
12,000 ft. The additional material, in excellent condition, is therefore of particular interest. It 
shows that hyacinthopa is remarkably widespread on Ruwenzori and occurs in the various floral 
zones from 4,500 to 14,500 ft. It shows also that the species is variable superficially. The wing 
expanse of the fresh material ranges from 19 mm. to 25 mm., compared with that of the type 
which measures 22 mm. Besides representing the norm in wing expanse the type specimen is 
remarkable in that it is also intermediate in the extent of white scaling in the distal area of the 
fore wing. In some of the additional specimens the white scaling is greatly reduced, while in 
others it is heavier and more extensive than in the type. 

Distribution: Uganda (Ruwenzori). 

Monopis liparota Meyrick 

Mouopis liparota Meyrick, 1920, Voyage de Ch. Alluaud et R. Jeannel en Afrique Oriental, 2, 
Microlepidoptera, p. 99. 

ruwenzori: Nyinabitaba, 8,650ft. (Fletcher), 3 q\ i?; Namwamba, 6,500 ft. (Edwards), 2 cm 
Namwamba, 10,200 ft. (Edwards), 1 6*- 
Distribution: Kenya. 

I am indebted to Dr. L. Gozmany for the identification of the above Ruwenzori material 
following his examination of the unique type of liparota in the Paris Museum. 



Monopis megalodelta Meyrick 
Moiwpis megalodelta Meyrick, 1908, Proc. zool. Soc. Lond., 1908 : 737. 
Uganda : Bundibugyo, 3,440 ft. (Fletcher), 1 $. 

Distribution: S. Nigeria, Sierra Leone and Natal. Bred from cocoons on Citrus leaf. 

Monopis sciagrapha sp.n. (Figures 57, 206, 207) 

o* 12-18 mm. Labial palpus yellowish white, overlaid with dark sepia exteriorly; terminal 
segment halt length of second, suffused with sepia, sometimes heavily, at base interiorly. Head 
light yellow. Thorax yellowish white, weakly irrorate with dark sepia laterally; tcgula dark 
sepia. Antenna sepia tinged greyish. Fore wing with veins 2 and 3 out of 4, 6 to apex, 7 and 8 
separate, 9, 10 and 11 separate, 11 from beyond middle of cell; yellowish white, variably 
suffused and irrorate with fuscous, suffusion strong along basal half of costa extending inwards to 
subcostal vein; a small dark sepia patch at extreme base of dorsum; a moderate subhyaline 
blackish sepia-edged depression in end of cell; a broad dark sepia apical streak extending 
into cilia, a similar but smaller and less conspicuous streak at tornus; cilia whitish yellow 
suffused fuscous. Hind wing with all veins separate; greyish bronze; cilia greyish. 
Male genitalia (Figs. 206, 207). 

ruwenzori: Ibanda, 4,700 ft., 4-12.ix.1952 (Fletcher), 2 including holotype, genitalia 
slide 9550; Ibanda 4,700 ft., 20-21. viii.1952 (Fletcher), 1 
Uganda: Bugoye, 4,500 ft., 5-10.ix.1952 (Fletcher), 1 A - 

Near M. coniodina Meyrick, from which it may be distinguished by the yellowish white 
coloration of the thorax, which in coniodina is dark fuscous, and by the presence of the dark 
sepia apical and tornal streaks. 

Tinea amphitrite Meyrick 
Tinea amphitrite Meyrick, 1932, Exot. Microlepidopt., 4 : 209. 

ruwenzori: Namwamba Valley, 10,200 ft. (Edwards), 1 6*; Namwamba Valley, Heath Zone, 
10,500-11,500 ft. (Edwards), 2 Namwamba Valley, 12-13,000 ft. (T. H. E.Jackson), 1 <J; 
Nyamaleju, 10,530 ft. (Fletcher), 7 Bigo, 11,400 ft. (Fletcher), 7 $\ Lake Bujuku, 13,050 ft. 
(Fletcher), 1 <J. 

This species has previously been known only from the type material : two males, not a male 
and female as stated by Meyrick in the original description, collected on Ruwenzori at 10,000 ft. 
The additional material reveals that considerable variation occurs in the coloration and fore 
wing markings in this species. In the typical (type) form the dark markings of the fore wing 
are strongly defined and contrast with the pale yellowish ground colour. Among the fresh 
material, six of the seven specimens from Bigo, 11,400 ft., are of the typical form, having the 
fore wing markings strong and well-defined, while the seventh specimen from tins locality has 
the fore wing strongly suffused with fuscous obliterating the pattern and making the wing an 
almost unicolorous brown in appearance. None of the specimens from other localities match 
the types exactly, and have the wing pattern variable and often reduced or partially obscured by 
fuscous suffusion. 

Distribution: Uganda (Ruwenzori). 


Ruwenzori Expedition, 1952. Volume II, Number 12 

Tinea othello Meyrick 
Tinea othello Meyrick, 1907, J. Bombay nat. Hist. Soc, 17 : 988. 
ruwbnzori: Ibanda, 4,700 ft. (Fletcher), 1 q*. 

Uganda: Semliki Forest, 2,850 ft. (Fletcher), 1 Bundibugyo, 3,440 ft. (Fletcher), 7 ex. 
Distribution: Widely distributed in Africa, Madagascar and India. 

Tinea allomella sp.n. (Figures 58, 59, 208-211) 

o* 13-18 mm. Labial palpus dark brown varying to greyish brown, a trace of yellowish suffusion 
at apex of terminal segment in some examples. Head dark brown with variable yellowish 
admixture. Thorax and tegula dark brown, individual scales on tegula and less frequently on 
thorax tipped with purplish. Antenna dark greyish brown. Fore wing dark brown varying to 
greyish brown in some examples, very weakly diffusedly irrorate with greyish; basal area often 
darkened with purplish which sometimes extends obliquely some distance along costal margin; 
a moderately prominent blackish stigma at end of cell; cilia yellowish orange. Hind wing greyish 
yellow, diffusedly overlaid with greyish brown except at apex and termen, darker towards base 
and along costa; sometimes a weak violaceous iridescence in examples having very dark colora- 
tion; cilia yellowish orange. Abdomen greyish brown. 

$ 10-12 mm. Brachypterous. Coloration and markings of fore wing basically as in male but 
paler and less intense. 

Male genitalia (Figs. 208, 209). 

Female genitalia (Figs. 210, 211). 

ruwenzori: Stuhlmann Pass, 13,500 ft., 27.viii.1952 (Fletcher), 20 <3, including holotype, 
genitalia slide 6530, 3 including allotype, genitalia slide 9584; Lake Bujuku, 13,050 ft., 
22-28.vii.1952 (Fletcher), 1 <$\ Namwamba Valley, 12-13,000 ft., xii. 1934-^1935 (Edwards), 
9 J'. Kumusong Valley, 12,000 ft., 23. xii. 1961-1.^1962 (Univ.-Rhod. Nyas.) 1 Nyamgasani 
Valley, 12-13,000 ft., xii. 193 4— i.193 5 (Buxton), 2 

Evans & Fletcher (1958 : v) comment on this species: "... found in numbers on the trunks 
of the Senccios, among the accumulated dead foliage, on which the larvae almost certainly feed". 

Closely related to T. luridula Meyrick, an alpine species from Abyssinia. The unique type of 
luriduh is in the British Museum and on re-examination during the present study was found 
to be a male and not female as stated by Meyrick in the original description. In general appear- 
ance it closely resembles allomella though somewhat paler in coloration; it differs most noticeably 
in having the head yellowish (pale ochraceous-orange) and in lacking the blackish stigma 
present on the fore wing in allomella. 

Tinea tolma sp.n. (Figures 60, 212, 213) 

6* 19 mm. Labial palpus dark fuscous, apical half of terminal segment pale yellow. Head pale 
chrome yellow. Antenna brownish grey. Thorax and tegula nutria colour, smooth, slightly 
shiny. Fore wing pale yellow, costa suffused nutria, dark towards base; an undefined tornal I 
spot of faint light grey suffusion, extending into cilia; cilia pale yellow. Hind wing brassy-grey; 
cilia whitish grey. 

Male genitalia (Figs. 212, 213). 

Microlepidoptera 119 

UGANDA : Bundibugyo, 3,440 ft., 22.v11i.-3.1x. 1952 (Fletcher), 1 holotype, genitalia slide 

Resembling the Abyssinian species T. nephelotorna Meyrick but with fore wing less brassy 
and hind wing a slightly darker shade of grey, sufficient to distinguish the two species superficially. 

Machaeropteris irritabilis Meyrick 
Machaeropteris irritabilis Meyrick, 1932, Exot. Microlepidopt., 4 : 329. 
UGANDA: Mbarara (Edwards), 1 3. 
Previously known only from the unique type male. 
Distribution: Uganda (Budongo Forest). 

Hapsifera horridella (Walker) 

Tinea horridella Walker, 1863, Cat. Lep. Ins. B.M., 2S : 474. 

ruwenzori: Ibanda, 4,700 ft. (Fletcher), 1 $. 
UGANDA: Bundibugyo, 3,440 ft. (Fletcher), 1 ?. 

Distribution: E. Africa, Congo and Natal. 

Hapsifera platyloxa Meyrick 
Hapsifera platyloxa Meyrick, 1930, Exot. Microlepidopt., 3 : 553. 

ruwenzori: Bwamba Pass (West side), 5,500-7,500 ft. (Edwards), 1 $. 
Distribution: Uganda. 


Narycia crocodilitis Meyrick 
Narycia crocodilitis Meyrick, 1930, Exot. Microlepidopt., 3 : 554. 

Uganda: Bundibugyo, 3,440 ft. (Fletcher), 2 Scmhki Forest, 2,850 ft. (Fletcher), 6 
Distribution: Uganda. 


Adela stenonipha sp.n. (Figures 61, 214) 

6* 12 mm. Head and thorax abraded, remaining scales indicate coloration may normally be 
blackish, with possibly yellowish admixture on front of head. Antennae broken, remaining basal 
section, including scape, purplish fuscous, a trace of overlaying whitish scaling on flagellum. 
Fore wing purplish fuscous with weak metallic golden sheen, purplish scaling very evident in 
discal area when viewed obliquely; a narrow well-defined slightly inward-oblique white fascia 
from costa at f, margins bordered with deep purplish suffusion to a depth about same as width 
of fascia, slightly greater on outer margin ; cilia greyish, mixed with purplish fuscous in basal 

120 Ruwenzori Expedition, 1952. Volume II, Number 12 

half. Hind wing dark grey, a weak purplish mixed with bronze sheen; cilia dark grey, a weak 
sub-basal line of deeper shade. Hind leg greyish, somewhat shining, tibia and first tarsal segment 
clothed above with long dark hair. 
Male genitalia (Fig. 214). 

ruwenzori: Nyinabitaba, 8,650 ft., 7-13.vii.1952 {Fletcher), 1 holotype, genitalia slide 

Near the Moroccan species A. collicolella Walsingham, but differing in the darker general 
coloration of the fore wing which in collicolella is shining bronzy metallic with a strong, distinc- 
tive golden sheen. 


Stigmella ruwenzoriensis sp.n. (Figures 62, 215) 

o* 8 mm. Head and collar (patagia) blackish brown; antemia greyish brown, antennal eyecaps 
cream colour. Thorax and tegula dark greyish brown with a very weak purplish sheen. Fore 
wing very elongate and narrow, in general appearance greyish brown, an indication of purplish 
sheen when viewed obliquely; individual scales light grey basally gradually becoming brown 
towards middle and dark apically, lighter coloured basal part of scales mostly concealed by 
overlapping scales; a moderately prominent cream colour tornal spot; cilia dark brown along 
costa and around apex, greyish with slight ochreous tinge along inner margin. Hind wing grey; 
cilia greyish, tinged ochreous along inner margin. Fore leg greyish brown, dorsal margin ot 
tibia suffused cream colour; middle and hind legs medium grey; hind tibia bearing 15 to 20 
well developed spines. Underside of fore wing greyish dark brown; hind wing dark grey. 

Male genitalia (Fig. 215). Tegumen broad proximally, tapering to a slender sparsely setose 
pseuduncus which terminates in a pointed tip bent ventrad. Uncus very distinctive, in the form 
of an inverted, quadrate, open horse-shoe with thick shoulders and lateral arms; arising 
ventro-medially is a short nail-headed projection. Valva broad basally, ventral margin irregularly 1 
serrulate; originating from inner surface near costa and close to arm of transtilla and projecting 
inwards is a style-like rod, conical, very slender, tapering gradually and with apex truncate. 
Ventral plate of vinculum rather short, breadth greater than width, caudal margin produced 
submedially to a pair of short triangular points, margin concave medially between these points 
and again immediately laterad of them; lateral arms of vinculum fused with large hood-shaped 
tegumen, forming with ventral plate a complete ring around the abdomen. Saccus extremely 
short, proximal margin concave, lateral angles obtusely rounded. Gnathus arms comparatively 
straight and narrow, merging medially to form a short, bluntly pointed tooth-like projection. 
Anellus developed distally in the form of two strongly sclerotized lobes (cornua), close to and 
laterad of aedeagus; juxta weakly sclerotized, deeply incised medially, laterally with two well 
sclerotized cornua which are slightly recurved and flattened basally. Aedeagus about four times 
as long as wide, slightly constricted near middle, distally with two lobes, one ventral and one 
dorsal; cornuti numerous, transparent or semitransparent, flat, spatulate or slender-triangulate. 

ruwenzori: Bigo, 11,400 ft., 20-22.vh.1952 {Fletcher), 1 <$, holotype, genitalia shde 8933. 

This species is apparently the first Nepticulid known from Ruwenzori, and has proved of 
considerable interest. It has been pronounced as certainly new by Mr. A. G. Carolsfeld-Krause 



of Copenhagen, Denmark, to whom I am indebted for guidance in compiling the foregoing 
description, notably of the genitalia characters, and for comments on the generic position of the 

S. ruwenzoriensis is closely related to the group of species from South Africa described in the 
genus Stigtnella by Vari (1955, Ann. Transvaal Mus., 22 : 331-336). It can be placed near S. 
oleivora Vari, from which it may be distinguished by the long and very narrow fore wing, and 
by the absence of markings on the fore wing except for the moderately prominent tornal spot. 
S. ruwenzoriensis is not typical of the genus Stigmella, and the present generic position should be 
regarded as provisional, having been adopted as a matter of expediency because the nearest 
apparent relatives have been assigned to that genus. The structure of the male genitalia shows 
certain characteristics of the palaearctic genus Fomoria Beirne, and it is likely that the generic 
afhnities of ruwenzoriensis and its congeners will prove to lie with that group. Certainly they 
belong in the dechtirioid section of the Nepticulidae with large hood-shaped tegumen. 





1. Trachybyrsis hypsitropha sp.n. holotype ^ (37 mm.) 5. Paraiuesiodcs aprcpta sp.n. liolotypc ' ' 17 2 

2. Capua spilonoma gitona subsp.n. holotype J mm., $ 19-24 mm.) 

18 mm.) 6. Niphothixa agelasta sp.n. liolotypc j (18111m.) 

3. Parapandcmis eustropha sp.n. holotype $ (image 7. Niphothixa ophina sp.n. liolotypc J (18 20111m 
reversed) (24-25 mm.) S. Tortrix edwardsi sp.n. holotype > (20 nun.) 

4. Parapandcmis orophila sp.n. holotvpe (j 1 (20 mm.) 


Ruwenzori Expedition, 1952. Volume II, Number 12 

10 14 

9. Tortrix stenophora sp.n. holotype <J (10-15111111.) 13. Epichoristodes heterotropha sp.n. holotype <J (18 
10. Metamesa octogona sp.n. holotype (16-19 mm.) mm.) 

[I. Epichoristodes panochra sp.n. paratype $ (14- 14. Eitcosma phaecchyta sp.n. holotype J 1 (22-23 mm.) 

16 mm.) 15. Notocelia scotodes sp.n. holotype <J (16 mm.) 

12. Epichoristodes atycta sp.n. paratype <§ (14-15 mm.) 16. Crimnologa fletcheri sp.n. allotype $ (26-33 mm.) 




17. En dothenia alpigena sp.n. holotypc ,£ (15-20 mm.) 21. Olethreutcs orestcra sp.n. holotypc j (24-27 mm.) 

18. Endothenia alpigena sp.n. allotype $ 22. Olethreutcs orestera sp.n. paracypc . (18-20 mm.) 

19. Endothenia alpigena sp.ri. paratype $ 23. Psaniathocrita dolonia sp.n. holotypc J (7-9 mm.) 

20. Epinotia penthfana sp.n. holotype (29-34 mm.) 24. Psaniathocrita doloma sp.n. allotype 


Ruwenzori Expedition, 1952. Volume II, Number 12 


25. Aristotelia cpacria sp.n. holotypc (16-19 mm.) 

26. Timyra floccula sp.n. holotype $ (22 mm.) 

27. Liniiiaecia pamphaea sp.n. holotype $ (15-16 


28. Mcridarchis mesosticha sp.n. holotypc (20 mm.) 


29. Carposina eusckema sp.n. holotypc $ (20-3 1 mm.) 

30. Carposina euschema sp.n. allotype $ 

31. Carposina mesophaea sp.n. holotype ^ (14 mm.) 

32. Carposina mesophaea sp.n. allotype § (16-18 mm.) 





33. Carposina poliophara sp.n. holotypc q (1S-20 37. Glyphipterix plagiophara sp.n. holotypc ?(n 
mm.) mm.) 

34. Choreutis pychnoinochla sp.n. holotypc $(15 mm.) 38. Elachista oritropha sp.n. holotypc * (10 11 mm.) 

35. Choreutis agelasta sp.n. holotype $ (13-15 mm.) 39. Elaclusta oritrapha sp.n. paratypc ,' 

36. Glyphipterix chalcotypa sp.n. paratypc o (12-16 40. Scythrh philoritcs sp.n. paratypc ,' (10 13 mm.) 


Ruwenzori Expedition, 1932. Volume II, Number 12 



41. Argyresthialamiella sp.n. holotype^ (11-12 mm.) 45. Metriochroa celidota sp.n. holotype g (7-9 mm.'. 

42. Argyresthia lamiella sp.n. allotype $ 46. Caloptilia pachyspila sp.n. holotype ^(9-12 mm. 

43. Ethmia phricotypa sp.n. holotype ^ (24 mm.) 47. Caloptilia pachyspila sp.n. paratype ^ 

44. Coleophora tacera sp.n. holotype ^ (11-15 mm.) 48. Caloptilia pachyspila sp.n. paratype 





49. Caloptilia janeae sp.n. holotypc $ (16-21 mm.) 

50. Caloptilia janeae sp.n. allotype $ 

51. Phyllocnistis loxositcha sp.n. paratypc J (7 mm.) 

52. Epernwnia oriplanta sp.n.holotype $ (13--7 mm.) 


53. Epcrmenia oriplanta sp.n. allotype 

54. Plutella syiimwrpha sp.n. holotypc > (10 mm. 

55. Acrolepia neplieloia sp.n. holotypc 3(11-12 mm. 

56. Tinissa polioplwsnia sp.n. holotypc (19 mm-) 


Ruwenzori Expedition, 1952. Volume II, Number 12 



57. Monopis sciagrapha sp.n. holotype $ (12-18 

58. Tinea allomella sp.n. holotype J 1 (13-18 

59. Tinea allomella sp.n. allotype $(10-12 mm.) 

60. Tinea tolma sp.n. holotype (19 mm.) 

61. Adela stenonipha sp.n. holotype $ (12 mm.) 

62. Stigmella ruwenzoriensis sp.n. holotype <J 
(8 mm.) 



63. Trachybyrsis hypsitropha sp.n. $ genitalia, 
ventral view, aedeagus in situ 

64. Capua spilonomagitona subsp.n. genitalia, 
ventral view, aedeagus removed 

65. Ibidem, aedeagus, lateral view 

66. Ibidem, enlargement of transtilla 

67. Parapandemis eustropha sp.n. 9 genitalia, 
ventral view 


68. Ibidem, ostium 

69. Ibidem, signuni 

70. Parapandemis orophila sp.n. J genitalia, 
ventral view, aedeagus in situ 

71. Paramesiodes aprepta sp.n. J 1 genitalia, 
ventral view, aedeagus in silu 

Ruwenzori Expedition, 19^2. Volume II, Number 12 



•• '. V"'* 





72. Paramesiodes aprcpta sp.n. ^ genitalia 

73. Paramesiodes aprepta sp.n. 5 genitalia, 
ventral view 

74. Ibidem, ostium 

75. Ibidem, signuni 


76. Niphothixa agelasta sp.n. ^ genitalia, 
ventral view, aedeagus in situ 

77. Niphothixa opliiua sp.n. ^ genitalia, ventral 
view, aedeagus in situ 

78. Niphothixa opliiua sp.n. ^ abdomen, lateral 
aspect, showing patches of specialised 
scales on sternum ot 1st segment 


85 (T 86 


79. Tortrix edwardsi sp.n. ^ genitalia, ventral 
view, aedeagus in situ 

80. Tortrix stenophora sp.n. ^ genitalia, ventral 
view, aedeagus removed 

81. Ibidem, aedeagus, lateral view 

82. Ibidem, coremata 

83. Tortrix stenophora sp.n. § genitalia, ventral 

f 1 G . 

84. Ibidem, ostium 

85. Ibidem, signum 

86. Mctamesia octogona sp.n. j genitalia, ventral 
view, aedeagus r'fi situ 

87. Epichoristodes panochra sp.n. _> genitalia, 
ventral view, aedeagus removed 

88. Ibidem, aedeagus, lateral view 


Ruwenzori Expedition, 1952. Volume II, Number 12 








m \ 










Epichoristodes atycta sp.n. ^ genitalia, ven- 
tral view, aedeagus removed 
Ibidem, aedeagus, lateral view 
Epichoristodes Iwterotropha sp.n. <J genitalia, 
ventral view, aedeagus removed 
Ibidem, aedeagus, lateral view 
Eucosma phaeochyta sp.n. g genitalia, 
ventral view, aedeagus in situ 


94. Notocelia scotodes sp.n. $ genitalia, ventral 
view, aedeagus in situ 

95. Crimnologa fletcheri sp.n. $ genitalia, ven- 
tral view, aedeagus in situ 

96. Crimnologa fletcheri sp.n. $ genitalia, ven- 
tral view 

97. Ibidem, ostium 















98. Endothenia alpigena sp.n. <J genitalia, 
ventral view, aedeagus removed 

99. Ibidem, aedeagus, lateral view 

100. Endothenia alpigena sp.n. § genitalia, 
ventral view 

101. Ibidem, ostium 

102. Ibidem, signum 


103. Epinotia penthrana sp.n. £ genitalia, ven- 
tral view, aedeagus in situ 

104. Epinotia penthrana sp.n. 9 genitalia, ven- 
tral view 

105. Ibidem, ostium 

106. Ibidem, sisna 


107. Olethreutes. orestera $ genitalia, ven- 111. 

tral view, aedcagus in situ 

toX. Olethreutes orestera sp.n. § genitalia, ven- 112. 

tral view 

109. Ibidem, ostium 113. 

no. Ibidem, signum 114. 

Psauiathocrita doloma sp.n. $ genitalia, 
ventral view, aedeagus in situ 
Psauiathocrita doloma sp.n. 9 genitalia, 
ventral view 
Ibidem, ostium 
Ibidem, signum 




115. Aristotelia cpacria sp.n. $ genitalia, lateral 
view, aedeagus in situ 

1 16. Aristotelia cpacria sp.n. abdomen, show- 
ing VHIth sternum and tergum 

117. Aristotelia cpacria sp.n. 9 genitalia, ventral 

118. Ibidem, ostium 

119. Ibidem, signum 


120. Timyra flocaila sp.n. j genitalia, ventral 

view, aedeagus removed 
[21. Ibidem, aedeagus, lateral view 

122. Liimiaecia patnphaea sp.n. j genitalia, 
lateral view, aedeagus in situ 

123. Liimiaecia pamphaca sp.n. J abdomen, 
showing Vlllth sternum and tcrquni 

Ruwenzori Expedition, 1952. Volume II, Number 12 

129 I 130 


124. Meridarchis mesosticha sp.n. genitalia, 
ventral view, aedeagus removed 

125. Ibidem, aedeagus, lateral view 

126. Carposina euschema sp.n. $ genitalia, ven- 
tral view, aedeagus removed 

127. Ibidem, aedeagus (subapical seta missing) 

128. Ibidem, aedeagus (subapical seta present) 

i3i : 132! 


129. Carposina euschema sp.n. 9 genitalia, 
ventral view 

130. Ibidem, ostium 

131. Carposina mesopbaea sp.n. $ genitalia, 
ventral view, aedeagus removed 

132. Ibidem, aedeagus, lateral view 




133. Carposina mesophaea sp.n. $ genitalia, 
ventral view 

134. Ibidem, ostium 

135. Ibidem, signa 

136. Carposina poliophara sp.n. q genitalia, 
ventral view, aedeagus removed 

137. Ibidem, aedeagus, lateral view 


138. Choreutis pychnomochla sp.n. $ genitalia, 
ventral view 

139. Ibidem, ostium 

140. Ibidem, sigiuim 

141. Choreutis agelasta sp.n. J 1 genitalia, ventral 
view, aedeagus removed 

142. Ibidem, aedeagus, lateral view 


Ruwenzori Expedition, 1952. Volume II, Number 12 


143 . Choreutis agelasta sp.n. ^ genitalia, ventral 

144. Ibidem, ostium 

145. Ibidem, signum 

146. Glyphiptcrix chalcotypa sp.n. genitalia, 
ventral view, aedeagus removed 

147. Ibidem, aedeagus, lateral view 


148. Glyphiptcrix chalcotpya sp.n. § genitalia, 
ventral view 

149. Ibidem, ostium 

150. Glyphiptcrix plagiophara sp.n. $ genitalia, 
ventral view, aedeagus removed 

151. Ibidem, aedeagus, lateral view 




152. Elachista oritropha sp.n. genitalia, ven- 

1 S7- 

Scythris philoritc'S sp.n. j 1 genitalia, ventral 

tral view, aedeagus removed 

view, tedeagus removed 

153. Ibidem, acdcagus, lateral view 


Ibidem, acdcagus, lateral view 

154. Elacliista oritropha sp.n. $ genitalia, ventral 

r S9- 

Scythris philoritcs sp.n. J 1 abdomen, show- 


ing hair-pencil arising from 1st segment 

t 55 . Ibidem, ostium 


Argyresthia lamiella sp.n. j genitalia, ven- 

156. Ibidem, signum 

tral view, acdcagus removed 


Ibidem, aedeagus, lateral view 


Kuwenzori Expedition, iqs2. Volume II, Number 12 

7 * V 







162. Argyresthia lamiella sp.n. ^ genitalia and 
coremata, ventral view, aedcagus re- 

163. Aroyresthia lamiella sp.n. $ abdomen, 
VHIth segment, showing V-shaped stern- 


164. Argyresthia lamiella sp.n. $ genitalia, ven- 
tral view 

165. Ibidem, ostium 

166. Ibidem, signum 

167. Ethmia phricotypa sp.n. £ genitalia, ven- 
tral view, aedeagus removed 

168. Ibidem, aedeagus, lateral view 



169. Coleophora tacera sp.n. $ genitalia, ventral 
view, aedeagus in situ 

170. Coleophora taccra sp.n. $ genitalia, ventral 

171. Ibidem, ostium 

172. Mctriochroa eclidota sp.n. £ genitalia, ven- 
tral view, aedeagus removed 


173. Ibidem, aedeagus, lateral view 

174. Metric .iron eclidota sp.n. 5 genitalia, sub- 
vent' il view 

175. Ibif'int, signum 

176. Caloptilia pachyspila sp.n. ^ genitalia, 

cntral view, aedeagus removed 

177. Ibidem, aedeagus, lateral view 


Ruwenzori Expedition, 1952. Volume II, Number 12 











[78. Caloptilia pachyspila sp.n., ^ genitalia, 

ventral view, aedcagus removed 
179. Ibidem, aedcagus, lateral view 
1 (So. Caloptilia pachyspila sp.n. ^ abdomen, 
showing coremata arising from Vllth 

181. Caloptilia janeae sp.n. genitalia, ventral 
view, acdeagus removed 


182. Ibidem, aedcagus, lateral view 
1 S3. Caloptilia janeae sp.n. $ abdomen, show- 
ing coremata arising from Vllth segment 

184. Caloptilia janeae sp.n. 9 genitalia, ventral 

185. Ibidem, ostium 

186. Ibidem, signa 


187. Phyllocnistis loxosticha sp.n. $ genitalia, 
including coremata, ventral view, aedea- 
gus removed 

188. Ibidem, aedeagus, lateral view 

189. Phyllocnists loxosticha sp.n. 9 genitalia, 
lateral view, ductus bursae and bursa 
copulatrix detached 

190. Ibidem, signa 


191. Epermenia criplanta sp.n. J 1 genitalia, ven- 
tral view, aedeagus removed 

192. Ibidem, aedeagus, lateral view 

193. Epermenia oriplanta sp.n. $ genitalia, ven- 
tral view 

194. Ibidem, ostium 

195. Ibidem, signum 


Kuwenzori Expedition, 1952. Volume II, Number 12 



196. Plutella symmorpha sp.n. J genitalia, ven- 
tral view, uncus and aedcagus removed 

197. Ibidem, uncus, lateral view 

198. Ibidem, acdeagus 

199. Acrolepia nephelota sp.n. genitalia, ven- 
tral view, aedcagus removed 

197 198 

20 1 202 


200. Ibidem, acdeagus, lateral view 

201. Acrolepia nephelota sp.n. $ genitalia, ven- 
tral view, bursa copulatrix detached 

202. Ibidem, bursa copulatrix 

203. Ibidem, ostium 




204. Tinissa poliophasma sp.n. § genitalia, ven- 
tral view 

205. Ibidem, ostium 

206. Monopis sciagrapha sp.n. £ genitalia, ven- 
tral view, aedeagus removed 

207. Ibidem, aedeagus, lateral view 

20S. Tinea allomeUa sp.n. 3 1 genitalia, ventral 

view, aedeagus removed 
209. Ibidem, aedeagus, lateral view 


Ruwenzori Expedition, 1952. Volume II, Number 12 


210. Tinea allomclla sp.n. $ genitalia, ventral 

2T 1. Ibidem, ostium 

2T2. Tinea tolmasp.n.$ genitalia, ventral view, 
aedcagus removed 


213. Ibidem, aedeagus, lateral view 

214. Adda stenonipha sp.n. $ genitalia, ventral 
view, aedeagus in situ 

215. Stigmella ruwenzoriensis sp.n. $ genitalia, 
ventral view, aedeagus in situ 


Riley Dunn &. Wilson LtJ